A Classification of the Living Sharks and Rays of the World

Introduction to the Third Edition

The following listing collects in a single reference the scientific nomenclature of the Recent

Elasmobranchii, listing genus-group and species-group names with authorities, and in the latter

group, type localities. In the case of the species recognized, geographical disrtibution and in

some cases bathymetric ranges are given and the nomenclature is arranged within a widely

accepted synthesis of higher groupings. No two systematists are in total agreement about the

placement of every single one of the 2000 or so basionyms here listed, so some compromise

has had to be reached in this respect. Where close relationships within a family or genus are

known, taxa are grouped together as such, otherwise they are listed in chronological order of

authorship date.

Common Abbreviations

FC (after species range) Australian Fishcode

Ra. Range

t/l type locality

Author Abbreviations

B. & S. or Big. & Schr. Bigelow & Schroeder

Bl. & Schn. Bloch & Schneider

Blkr. Bleeker

Bonn. Bonnaterre

Dum. Dumeril

Fwlr Fowler

Gar. Garman

M.& H. Mueller & Henle

Raf. Rafinesque

GENERAL CLASSIFICATION

Superorder: HEXANCHOIDEA p. 4

Order Hexanchiformes p. 4

Superorder: HETERODONTOIDEA p. 5

Order Heterodontiformes p. 5

Superorder: SQUALOIDEA p. 7

Order Squaliformes p. 7

Order Pristiophoriformes p. 24

Order Squatiniformes p. 25

Superorder: GALEOIDEA p. 28

Order Orectolobiformes p. 28

Order Lamniformes p. 36

Order Carcharhiniformes p. 45

Superorder: BATOIDEA p. 94

Order Pristiformes p. 94

Order Rhinobatiformes p. 96

Order Torpediniformes p. 107

Order Rajiformes p. 116

Order Myliobatiformes p. 171

Chlamydoselachus GARMAN 1884, type Chlamydoselachus anguineus GARMAN 1884

Chlamydoselache GUNTHER 1887 (Emendation)

Chlamydoselachoides FOWLER 1947 type Chlamydoselachus lawleyi DAVIS

(Fossil )

Chlamydoselachus anguineus GARMAN 1884 t/l; Japanese Seas.

Range; Japan; Norway to Portugal; California.

FC 00 006001

Chlamydoselachus sp. EBERT t/l; S. Africa.

Hexanchus RAFINESQUE 1810, type Squalus griseus BONNATERRE 1788

Squalus (Monopterhinus) BLAINVILLE 1816 (part), type Squalus griseus BONNATERRE 1788

Notidanus CUVIER 1817 (part), type Squalus griseus BONNATERRE 1788

Hexanchias SWAINSON 1838 (Emendation)

Hexancus AGASSIZ 1846 ( ? lapsus), type Squalus griseus BONNATERRE 1788

Vacca SCILLA 1759

Le Griset BROUSSONET 1780 ( Mediterranean)

Squalus griseus BONNATERRE 1788 t/l; Mediterranean

Squalo SPALLANZANI 1793 ( Italy)

Le Squale Griset LACEPEDE 1798

Squalus vacca BLOCH & SCHNEIDER 1801

Hexanchus griseus RAFINESQUE 1810 t/l; Sicily

Squalus (Monopterhinus) colombinus BLAINVILLE 1816

Notidanus monge RISSO 1826 t/l; Nice, France

Notidanus sp. dubia POEY 1865 t/l; Cuba

Hexanchus corinus JORDAN & GILBERT 1880 t/l; Washington; Monterey Bay

Notidanus vulgaris PEREZ-CANTO 1886 t/l; Chile.

Hexanchus griseus australis De BUEN 1960

Range;E. Atlantic & Med. south to Cape Verde Is., Southern Africa but not definitely Cape region. New England to Northern Gulf of Mexico. Alaska to Baja California. Hawaii. Japan. Dubious from Indonesia. Chile. N. Argentina & S.Brazil. S.E. Aus. N.Z.

FC 00 005005

Hexanchus griseus nakamurai TENG 1962 t/l; Taiwan

Hexanchus vitulus SPRINGER & WALLER 1969 t/l; Bimini, Bahamas.

Range; Bahamas; Florida; East Africa; Madagascar;Philippines.

FC 00 005004

Heptranchias RAF. 1810, type Squalus cinereus GMELIN 1789

? Squalus (Monopterhinus) BLAINVILLE 1816 (part) (? M. ciliaris BLAINVILLE 1816)

Notidanus CUVIER 1829 ( part )

Heptanchus MULLER& HENLE 1841 (emendation) type Squalus cinereus GMELIN 1789

Aellopos AGASSIZ 1843 type Aellopos wagneri AGASSIZ (fossil)

Heptranchus GRAY 1851 (emendation)

Notidanion JORDAN 1923 type Notidanus primigenius AGASSIZ (fossil)

Le Perlon BROUSSONET 1780

Squalus perlo BONN. 1788 t/l; Mediterranean

Squalus cinereus GMELIN 1789 t/l; Mediterranean

Le Squale perlon LACEPEDE 1798

Heptranchias cinereus RAF. 1810 t/l; Sicily

? Squalus (Monopterhinus) ciliaris BLAINVILLE 1816 (n.nud.)

Notidanus monge var. griseus (RISSO 1826) BONAPARTE 1846(name in synonymy)

Notidanus monge var. albescens (RISSO 1826) BONAPARTE 1846 (name in synonymy)

Notidanus monge var. rubescens (RISSO 1826) BONAPARTE 1846 (name in synonymy)

Heptrancus angio COSTA 1854 t/l; Naples

Notidanus (Heptrancus) cinereus var. pristiurus (var. aetatis) BELLOTTI 1877 t/l; Med.

Heptranchias deani JORDAN & STARKS 1901 t/l; Misaki, Japan.

Heptranchias dakini WHITLEY 1931 t/l; Australia.

Range; Worldwide in tropical & subtropical seas.

FC 00 005001

Notorynchus AYRES 1855, type Notorynchus maculatus AYRES 1855 ( = Squalus cepedianus PERON 1807) N. CEPEDIANUS (Peron ) 1807 Broadhead Sevengill Shark

Squalus cepedianus PERON 1807 t/l; Adventure Bay, Tasmania.

? Notidanus indicus AGASSIZ 1835 t/l; East Indies. ( teeth )

Notorynchus maculatus AYRES 1855 t/l; California

Notorhynchus borealis GILL 1864 t/l; Oregon (jaws)

Heptranchias pectorosus GARMAN 1884 t/l; Patagonia

Notidanus ferox PEREZ-CANTO 1886 t/l; Chile

Heptranchias haswelli OGILBY 1897 t/l;" Cape of Good Hope"(?Australia)

Notidanus medinae PHILIPPI 1901 t/l; Cartagena, Chile

Notidanus wolniczkyi PHILIPPI 1901 t/l; Coquimba, Chile

Notidanus wolniczkii PHILIPPI 1901(emendation)

Heptranchias spilotus LAHILLE 1913

Notorhynchus ocellatus DEVINCENZI 1920 t/l; Uruguay

Notorhynchus macdonaldi WHITLEY 1931 t/l; NSW., Aus.

Range;S.Aus.; Tasmania; S. Alaska to Baja Caifornia;Peru; Chile; Argentina; S. Brazil;Japan & Eastern China; S. Africa . Indian records dubious.

FC 00 005002

Species dubiae :-

Squalus platycephalus TENORE 1809 (nom. nud.) t/l; off Naples, Italy

Notidanus sp. CUVIER 1829

Cestracion SCHAEFFER 1760 (Inadmissible work) Not binomial

Squalus (Heterodontus) BLAINVILLE 1816, type Squalus philippi BLOCH & SCHNEIDER 1801. Monotypic.

Cestracion CUVIER 1817, type Squalus philippi BLOCH & SCHNEIDER 1801.

Cestracion OKEN 1817 (Inadmissible work) Refers to Schaeffer 1760, also inadmissible.

non Cestracion KLEIN 1742, KLEIN 1776, WALBAUM 1792, GILL 1862 and GARMAN 1913 which equal Sphyrna RAFINESQUE 1810 ( q.v.)

? Drepanophorus EGERTON 1872, type Acrodus rugosus AGASSIZ (fossil)

Port Jackson Shark PHILLIP 1789 ( Port Jackson,Botany Bay, NSW., Australia )

Squalus portus jacksoni MEYER 1793 (On Phillip) t/l; Port Jackson,Botany Bay, NSW., Australia

Le squale Philipp LACEPEDE 1798

Squalus jacksoni SUCKOW 1799 t/l; Port Jackson,Botany Bay, NSW., Australia

Squalus philippi BLOCH & SCHNEIDER 1801 (On Phillip) t/l; Port Jackson,Botany Bay, NSW., Australia.

Squalus philippinus SHAW 1804 (On Phillip) t/l; Port Jackson,Botany Bay, NSW., Australia

? Heterodontus phillipi var. japonicus DUMERIL 1865 (part) t/l; Australia

Cestracion heterodontus SHERRARD 1896 t/l;Hobson's Bay,Victoria.

Cestracion phillippi REGAN 1908 (lapsus)

Heterodontus bonae-spei OGILBY 1908 t/l; "Table Bay,S. Africa" in error.

Range; Australia; N.Z.

FC 00 007001

Gyropleurodus GILL 1862, type Cestracion francisci GIRARD 1854. Monotypic.

Tropidodus GILL 1862, type Cestracion pantherinus VALENCIENNES 1846 (1855) = Cestracion quoyi FREMINVILLE 1840.

Cestracion quoyi FREMINVILLE 1840 t/l; Galapagos Is.

Cestracion pantherinus VALENCIENNES 1845

Gyropleurodus peruanus EVERMANN & RADCLIFFE 1917 t/l; Lobos de Tierra I., Peru

Range; Galapagos Is.; Peru.

Cestracion francisci GIRARD 1854 t/l; Monterey Bay, California.

Heterodontus californicus HERALD 1961 (error for francisci)

Range; Central California to Gulf of California; ?Ecuador.

Gyropleurodus ramalheira J.L.B.SMITH 1949 t/l; Imhambane,Mozambique.

Range;S. Mozambique and Natal and off S.E.Arabia.

Heterodontus mexicanus TAYLOR & CASTRO-AGUIRRE 1972 t/l; Ceiro Colorado,Sonora,Mexico.

Range; Gulf of California, probably south to Peru.

Molochophrys WHITLEY 1931, type Cestracion galeatus GUNTHER 1870

Cestracion galeatus GUNTHER 1870 t/l;N.S.W., Australia

Range; N.S.W. and Queensland, Australia.

FC 00 007003

Centracion GRAY 1831 (emended variant spelling of Cestracion) , type Centracion zebra GRAY 1831. Monotypic.

Wuia FOWLER 1934, type Centracion zebra GRAY 1831.

Centracion zebra GRAY 1831 t/l; Swatow, China.

? Cestracion philippi var. japonicus DUMERIL 1865 (part) (Australia)

Cestracion phillipi var. zebra MARTENS 1876 (Nagasaki)

Cestracion amboinensis REGAN 1906 t/l; Amboina

Range; Southern Japan;Eastern China to Indo-China. Celebes. N. Australia.

DC 00 007002

? Heterodontus philippi var. japonicus DUMERIL 1865 (part) (Japan)

Cestracion philippi MULLER & HENLE 1841 (non BLOCH & SCHNEIDER 1801) (Japan) Not the "New Holland" example, which is portusjacksoni

Heterodontus phillippi BREVOORT 1856 (Simola)

Cestracion japonicus MACLAY & MACLEAY 1884 t/l; Tokyo, Japan.

Range; Japan; Korea; China.

Heterodontus sp. MEE Oman Oman Bullhead Shark

Squalus LINN. 1758 type Squalus acanthias LINN.1758

Squallus SCOPOLI 1777 type Squalus acanthias LINN.1758

Squalis RADERMAKER 1779

Squalus (Acanthorhinus) BLAINVILLE 1816 (part) incl. Squalus acanthias LINN.1758

Spinax CUVIER 1817 (part)

Acanthias LEACH 1818 (nom. nud.)

Acanthias RISSO 1826 type Acanthias vulgaris 1826 = Squalus acanthias LINN. 1758

Carcharias GISTEL 1848( non RAF.) to replace Acanthias RISSO 1826

Centrophorides DAVIS 1887 type Centrophorides latidens DAVIS 1887( fossil )

Squalus (Flakeus) WHITLEY 1939 type Squalus megalops MACLEAY

Squalus (Koinga) WHITLEY 1939 type Squalus griffini PHILLIPS ( = Squalus mitsukurii )

Akanthias galeos ARISTOTLE Historia Animalium VI c. 10, IX c. 37

Mustelus spinax BELON 1553 (not LINNAEUS 1758 = Etmopterus)

Galeus acanthias ROND. 1554

Stachdhund GESNER 1558

Squalus sp. no. 1 ARTEDI 1738

Squalus sp. no. 3 ARTEDI 1738

Squalus acanthias LINNAEUS 1754 Mus. Idol. Frid. p. 53

Squalus acanthias LINNAEUS 1758 ("Oceano Europaeo")

Picked Dog BROOKES 1763

Spinax DUHAMEL 1769-82 (inadmissible work)

Piked Dog BROOKES 1772

Piked Dogfish PENNANT 1776 t/l; Cornwall, England.

L'Aiguillet BROUSSONET 1780 (Mediterranean, Atlantic & Scotland)

Squalus spinax OLAVIUS 1780 (Iceland)

Squalus fernandinus MOLINA 1782 t/l;Chile

Squale aiguillet LACEPEDE 1798

Acanthorhinus ferdinandinus BLAINVILLE 1816 (lapsus for fernandinus)

Acanthias antiquorum LEACH 1818 (n. nud.)

Acanthias vulgaris RISSO 1826 t/l; Nice

Acanthias americanus STORER 1846 New York to Labrador

Spinax fernandezianus GUICHENOT 1848 t/l; Chile

Spinax mediterraneus GISTEL 1848 t/l; Mediterranean

Spinax (Acanthias) suckleyi GIRARD 1854 t/l; Puget Sound, Washington

Acanthias sucklii GIRARD 1858 (emendation)

Acanthias linnei MALM 1877 t/l; Goteberg, Sweden

Acanthias lebruni VAILLANT 1888 t/l; Straits of Magellan

Acanthias commun NAVARETTE 1898

Squalus mitsukurii TANAKA 1917 (non JORD. & FWLR. 1903)

Squalus wakiyae TANAKA 1918 t/l; Japan

Squalus kirki PHILLIPPS 1931 t/l; New Zealand

Squalus whitleyi PHILLIPPS 1931 t/l; Victoria, Australia

Squalus barbouri HOWELL-RIVERO 1936 t/l; Havana, Cuba

Squalus acanthias africana MYAGKOV & KONDYURIN 1986 (southeast Atlantic)

Spurdog; Piked Dogfish; Spiny Dog; Speerdog; Bone Dog; Skittledog;

Hoe; Cu-maire (= Sea Dog); Gobag; Ci-pigog; Doorhael.

Ra: Iceland and Norway to Mediterranean; Labrador to Cuba; W. coast of South Africa; Washington to Alaska round to Japan; temperate Australia; New Zealand; Chile to Straits of Magellan. Temperate seas worldwide. FC 00 020008

Acanthias blainville RISSO 1826 t/l;Nice, France

Spinax blainvillii BONAPARTE 1834 (emendation) t/l; Italy

Squalus blainvillei GRIFFINI 1903 (emendation)

Ra: Mediterranean and Black Sea; Canaries; Portugal; tropical & subtropical east Atlantic.

Acanthias megalops MACLEAY 188 t/l; Pt. Jackson; NSW

? Squalus acutipinnis REGAN 1908 t/l; Table Bay, South Africa

Squalus tasmaniensis HOWELL-RIVERO 1936 t/l;Tasmania

Ra: ; Queensland to S. & W. Aus.; Tasmania; Japan. ( ?South Africa)

FC 00 020006

Squalus mitsukurii JORDAN & FOWLER t/l; 1903 t/l; Misaki, Japan

Squalus philippinus SMITH & RADCLIFFE 1912 (non SHAW 1804) t/l; off Luzon, Philippines.

Squalus montalbani WHITLEY 1931 (nom. nov. for philippinus S & R, preocc.)

? Squalus griffini PHILLIPPS 1931 t/l; Hauraki Gulf, New Zealand

Ra: Japan to Philippines. Queensland to N.W. Australia via southern Australia & Tasmania. ?N.Z.

FC 00 020007

Squalus japonicus ISHIKAWA 1908 t/l; Kagoshima, Japan

Ra: Japan

Squalus brevirostris TANAKA 1912 t/l; Tokyo market (from Shimonoseki)

Ra; Japan.

Galludo PARRA 1787

Squalus cubensis HOWELL-RIVERO 1936 t/l; Havana, Cuba

Ra: off Havana and Matanzas, Cuba

Squalus asper MERRETT 1973 t/l; Aldabra I.,S.W.Indian Ocean.

Ra: Western Indian Ocean; Gulf of Mexico; widespread in deep water

Squalus melanurus FOURMANOIR & RIVATON 1979 t/l;New Caledonia

Ra: New Hebrides; New Caledonia

Squalus rancureli FOURMANOIR et. al. 1979 t/l; New Hebrides.

Range; New Hebrides;New Caledonia.

S. ACUTIROSTRIS Chu,Meng & Li 1984 Acutenose Spurdog

Squalus acutirostris CHU, MENG & LI 1984 t/l; South China Sea.

Range; South China Sea.

Squalus probatovi MYAGKOV & KONDYURIN 1986 t/l; S.E. Atlantic.

Range; S.E. Atlantic.

Range; off Queensland between Cairns& Rockhampton

FC 00 020037

Range;Upper Continental Slope off Queensland

FC 00 020038

Range;Slope off Western Australia

FC 00 020018

Range;Upper slope off NW Australia

FC 00 020039

Range;Slope off Western Australia Scott Reef to Perth.

FC 00 020040

Range;Slope, Queensland. Cape York to Rockhampton.

FC 00 020041

? Squalus (Acanthorhinus) BLAINVILLE 1816 (part) incl. infernus = uyato.

Centrophorus MULLER & HENLE 1837, type Squalus granulosus BLOCH & SCHNEIDER 1801

Entoxychirus GILL 1862, type Squalus uyato RAFINESQUE 1810

Atractophorus GILCHRIST 1922, type Atractophorus armatus GILCHRIST 1922

Gaboa WHITLEY 1940, type Centrophorus harrissoni McCULLOCH 1915

Squalus granulosus BLOCH & SCHNEIDER 1801 No type loc. Holotype apparently lost.

Centrophorus bragancae REGAN 1906 t/l; Cecimbra, Portugal.

Centrophorus tessellatus GARMAN 1906 t/l; 35 N., 139 30 E.,east of Japan in 400 fthms.

Centrophorus atromarginatus GARMAN 1913 t/l; Suruga Bay, Japan.

Centrophorus machequensis MAUL 1955 t/l; Madeira

Range; Bay of Biscay, Mediterranean, Madeira and Eastern Atlantic to Zaire; Natal to Northern Madagascar; Southern Japan to South China Sea; S. Australia; Hawaii; Gulf of Mexico.

Australian FC 00 020023

C. UYATO (Rafinesque) 1810 Little Gulper Shark; Harrisson's Deepsea Dogfish

Squalus uyato RAFINESQUE 1810 t/l; Sicily

Squalus uyatus RAFINESQUE 1810

? Squalus (Acanthorhinus) infernus BLAINVILLE 1816 (n.nud.)

Squalus infernus BLAINVILLE 1825 t/l; Mediterranean

Acanthias nigrescens NARDO 1860 t/l; Italy

Centrophorus harrissoni McCULLOCH 1915 t/l; near Cabo I., Victoria, Australia

Centrophorus ujato TORTONESE 1956 (lapsus)

Centrophorus niaukang TENG 1959 t/l; Taiwan.

Centrophorus armatus barbatus TENG 1962 t/l; East China Sea.

Range; Western Mediterranean to Namibia; Mozambique; S.E. Australia; East China Sea.

Australian FC 00 020010 (harrissoni) Australian FC 00 020011 (uyato)

Centrophorus moluccensis BLEEKER 1860 t/l; Amboina, Moluccas.

Centrophorus scalpratus McCULLOCH 1915 t/l; Victoria, Australia.

Atractophorus armatus GILCHRIST 1922 t/l; Natal, S. Africa.

Range; ;Moluccas; Southern Africa from Natal to Mozambique. S. E. Australia. New Hebrides. New Caledonia.

FC 00 020001

Centrophorus lusitanicus BOCAGE & CAPELLO 1864 t/l; Portugal

? Centrophorus ferrugineus CHU, MENG & HU 1982 t/l; South China Sea.

Range; Eastern Atlantic; Southeast Africa; Taiwan; ? South China Sea

Lepidorhinus BONAPARTE 1838, type Squalus squamosus BONNATERRE 1788

Machephilus JOHNSON 1867, type Machephilus dumerili JOHNSON 1867

Somnispinax WHITLEY 1940, type Centrophorus nilssoni THOMPSON 1930.

Enchiridiodon J.L.B. SMITH 1967, type Enchiridiodon hendersoni J.L.B. SMITH 1967

L' ecailleux BROUSSONET 1780

Squalus squamosus BONNATERRE 1788

Machephilus dumerili JOHNSON 1867

Centrophorus foliaceus GUNTHER 1877 t/l; Inosima, Japan.

Centrophorus acus GARMAN 1906 t/l; Japan.

Centrophorus steindachneri PIETSCHMANN 1907 t/l; Japan

? Lepidorhinus kinbei TANAKA (?date) ?Japan.

? Centrophorus drygalskii ENGELHARDT 1912 t/l; Japan

Centroscymnus fuscus GILCHRIST & VON BONDE 1924 t/l; S. Africa.

Centrophorus nilssoni THOMPSON 1930 t/l; off Kaikoura, N.Z.

Enchiridiodon hendersoni J.L.B. SMITH 1967 t/l; Algoa Bay, South Africa.

Range; Eastern Atlantic from Iceland to Senegambia; S. Africa; S. Japan; Philippines; S.E.Australia; New Zealand; North of Madagascar. Northern Gulf of Mexico

FC 00 020009

Centrophorus squaloideus ENGELHARDT 1913 (n.nud.)

Pseudocentrophorus CHU, MENG & LU 1981, gen. nov. type Pseudocentrophorus isodon CHU, MENG & LU 1981 sp. nov. Blackfin Gulper Shark t/l; South China Sea.

Centrophorus robustus DENG, XIONG & ZHAN 1985 t/l; East China Sea.

Centrophorus sp. SERET New Caledonia.

Acanthidium LOWE 1839 (In part) included A. calceus LOWE.

Deania JORDAN & SNYDER 1902 type Deania eglanteria JORDAN & SNYDER 1902

Deaniops WHITLEY 1932, type Acanthidium quadrispinosum McCULLOCH 1915

Acanthidium calceum LOWE 1839 t/l; Madeira (No type material)

Centrophorus crepidalbus BOCAGE & CAPELLO 1864 t/l; Portugal.

Deania eglantina JORDAN & SNYDER 1902 t/l; Totomi Bay, Japan.

Acanthidium rostratum GARMAN 1906 t/l; Suruga Gulf, Japan.

Acanthidium aciculatum GARMAN 1906 t/l; Sagami Bay, Japan.

Centrophorus kaikourae WHITLEY 1934 t/l; Off Kaikoura, New Zealand

Deania calcea BIGELOW & SCHROEDER 1957

Range; Noth Eastern Atlantic; Namibia to Algoa Bay,S.Africa;Japan; New Zealand.

FC 00 020003

Acanthidium hystricosum GARMAN 1906 t/l; Sagami Bay, Japan.

Deania mauli CADENAT 1983 t/l; Madeira.

Range; Japan;Madeira.

Acanthidium quadrispinosum McCULLOCH 1915 t/l; Gabo I.,Gt. Australian Bight and Victoria, Australia.

Range; S.Australia;Victoria, Aus.; South Africa from Namibia to S.Mozambique.

FC 00 020004

Nasisqualus SMITH & RADCLIFFE 1912, type Nasisqualus profundorum SMITH & RADCLIFFE 1912.

Nasisqualus profundorum SMITH & RADCLIFFE 1912 t/l; Between Leyte & Mindanao, Philippines.736 - 976 fthms.

Acanthidium natalense GILCHRIST 1922 t/l; Natal, S.Africa.

Deania elegans SPRINGER 1959 t/l; Cape Fear,North Carolina.

Deania cremouxi CADENAT 1960 t/l; Senegal. W.Africa.

Range; Both sides of the Atlantic & Western Pacific Oceans.

Cirrhigaleus TANAKA 1912, type Cirrhigaleus barbifer TANAKA 1912

Phaenopogon HERRE 1935, type Phaenopogon barbulifer HERRE 1935

Cirrhigaleus barbifer TANAKA 1912 t/l; Tokyo Market. From S.E.Honshu, Japan.

Phaenopogon barbulifer HERRE 1935 t/l; Japan.

Range; Japan; S.E.Australia; N.Z.

FC 00 020026

Etmopterus RAFINESQUE 1810, type Etmopterus aculeatus RAF.1810 = Squalus spinax LINN. 1758

Squalus (Spinax) CUVIER 1817, type Squalus spinax LINN. 1758

Acanthias RISSO 1826 (part ) incl. spinax.

Centrina LOWE 1833 (part),type Centrina nigra LOWE = Squalus spinax LINN.

Galeus acanthias seu Spinax fuscus : Genuae, Sagree WILLUGHBY 1686

Squalus sp. 4 (pinna ani carens; naribus in extremo rostro) ARTEDI 1738

Squalus spinax LINNAEUS 1746 Fauna Svec. p.107

Squalus spinax LINNAEUS 1758 Syst. nat. 1.p. 233 t/l;Oceano Europaeo

Squalus niger GUNNERUS 1762 t/l; Norway (Trondheim)

Le blaataske ASCANIUS 1777

Le sagre BROUSSONET 1780

Le squale sagre LACEPEDE 1798

Etmopterus aculeatus RAFINESQUE 1810 t/l; Sicily

Spinax gunneri REINHARDT 1828 t/l; Iceland

? Spinax vitalinus DE LA PYLAIE 1835

Spinax linnei MALM 1877 t/l; Sweden

Range; North East Atlantic & Med.

Etmopterus princeps COLLETT 1904 t/l; Faroe Channel, 750 - 1200 metres

Range; N.E. Atlantic.

Spinax unicolor ENGELHARDT 1912 t/l; Honshu, Japan.

Range; Honshu, Japan.

Etmopterus baxteri GARRICK 1957 t/l;South of Kaiakoura, N.Z.

Range; N.Z.

Spinax hillianus POEY 1861 t/l; Havana, Cuba.

Etmopterus polli BIGELOW, SCHROEDER & SPRINGER 1953 t/l; N. Angola,West Africa.

Etmopterus gracilispinis KREFFT 1968 t/l; Continental shelf off Uruguay

Range; Western Atlantic from Chesapeake Bay to Uruguay. Eastern South Atlantic.

Etmopterus virens BIGELOW, SCHROEDER & SPRINGER 1953 t/l; Northern Gulf of Mexico.

Range; Northern Gulf of Mexico.

Etmopterus schultzi BIGELOW, SCHROEDER & SPRINGER 1953 t/l;Northern Gulf of Mexico.

Range; Northern Gulf of Mexico.

E. ROBINSI Schofield & Burgess 1997

Etmopterus robinsi SCHOFIELD & BURGESS 1997 t/l; N.W. Providence Channel, Bahamas.

Range; Bahamas.

Etmopterus granulosus group

Spinax granulosus GUNTHER 1880 t/l; Southwest South America

Range; Southern Argentina & Chile; Tasmania

FC 00 020021

Etmopterus lucifer JORDAN & SNYDER 1902 t/l; Misaki, Japan.

Etmopterus brachyurus SMITH & RADCLIFFE 1912 t/l; 6 N., 120 E.,off Jolo, Philippines.

Acanthidium molleri WHITLEY 1939 t/l; N.S.W., Australia.

Etmopterus abernethyi GARRICK 1957 t/l; New Zealand.

Range; Japan; Philippines; Southern half of Australia; N.Z.

Australian fishcodes 00 020005 (lucifer) ; 00 020032 (brachyurus) ; 00 020033 (molleri)

Etmopterus villosus GILBERT 1905 t/l; Molokai, Hawaii.

Etmopterus bullisi BIGELOW & SCHROEDER 1957 t/l; N.E.coast of Florida.

Range; Hawaiian Is. Western N.Atlantic, N. Carolina to Caribbaean

Etmopterus decacuspidatus CHAN 1966 t/l; Hainan, China.

Range; Hainan

Etmopterus sentosus group

Etmopterus sentosus BASS 1976 t/l;Natal Basin;Bazaruto I., Mozambique

Range; Natal Basin;Bazaruto I., Mozambique

Acanthidium LOWE 1839, type Acanthidium pusillum LOWE 1839

Centrina nigra LOWE 1843 (non CUVIER) t/l; Madeira

Acanthidium pusillum LOWE 1839 t/l; Madeira

? Etmopterus frontimaculatus PIETSCHMANN 1907 t/l; Sagami Sea, Japan

Range; N.E. Atlantic; Japan; Tasmania; Central N.S.W.,Aus.

FC 00 020015

Etmopterus bigelowi SHIRAI & TACHIKAWA 1993 t/l; Angola

Range ; off Angola

Etmopterus carteri SPRINGER & BURGESS 1985 t/l; Caribbaean coast of Colombia.

Etmopterus perryi SPRINGER & BURGESS 1985 t/l; Caribbaean coast of Colombia.

Etmopterus schmidti DOLGANEV 1986 t/l; N.W.Pacific.

Etmopterus tasmaniensis MYAGKOV & PAVLOV 1986 t/l;Tasmania

Etmopterus splendidus YANO 1987 t/l; Japan.

Etmopterus compagnoi FRICKE & KOCH 1990 t/l; Cape Basin, South Africa.

Etmopterus litvinovi PARIN & KOTLYAR 1990 t/l; Eastern Pacific

Etmopterus pycnolepis PARIN & KOTLYAR 1990 t/l; Eastern Pacific

Etmopterus sp. A LAST & STEVENS 1993 Abrolhos Is. to Sunbury, W. Aus.;Sydney, N.S.W. to Tasmania. ? Synonym of E. bigelowi SHIRAI & TACHIKAWA. Australian fishcode 00 020027

Etmopterus sp. B LAST & STEVENS 1993 Perth. W.Aus. to Taree, NSW. FC 00 020022

Etmopterus sp. C LAST & STEVENS 1993 off Broome, Western Aus. ? Synonym of E. splendidus YANO 1987 Australian fishcode 00 020028

Etmopterus sp. D LAST & STEVENS 1993 Cairns, N. Queensland. Australian fishcode 00 020029

Etmopterus sp. E LAST & STEVENS 1993 Rowley Shoals & Ashmore Reef, W. Aus. Australian fishcode 00 020030

Etmopterus sp. F LAST & STEVENS 1993 Between Cairns & Rockhampton, North Queensland.

Australian fishcode 00 020031

Etmopterus sp. COMPAGNO Guadalupe, Lesser Antilles.

Etmopterus sp. COMPAGNO Chile

Etmopterus sp. cf. princeps COMPAGNO Inaccessible Island

Etmopterus sp. COMPAGNO & EBERT Sculpted Lantern Shark S. Africa.

Etmopterus sp. cf. granulosus or baxteri COMPAGNO & EBERT Southern Africa.

Genus CENTROSCYLLIUM Muller & Henle 1841

Subgenus CENTROSCYLLIUM Muller & Henle 1841

Centroscyllium MULLER & HENLE 1841, type Spinax fabricii REINHARDT 1825

Squalus acanthias FABRICI 1780 (non LINN. 1758) t/l; Julianahaab, West Greenland.

Spinax fabricii REINHARDT 1825 t/l; West Greenland

Range; North Atlantic from Davis Strait, West Greenland; Iceland;Faroes;Newfoundland. Below 150 fthms.

Centroscyllium granulatum GUNTHER 1887 t/l;Port Stanley, Falkland Is. in 448 metres.

Range;Off Southern South America.

Centroscyllium nigrum GARMAN 1899 t/l; Between 6&7 degrees N. and 81 & 82 degrees W.,Eastern Pacific, off Panama in 546 - 555 fthms.

Centroscyllium ruscosum GILBERT 1905 t/l; Kauai, Hawaiian Is.

Range; Panama;Galapagos Is.;Cocos I.;Peru;Hawaiaan Is.

Centroscyllium ritteri JORDAN & FOWLER 1903 t/l; Misaki, Japan.

Range; Japan.

Centroscyllium kamoharai ABE 1966 t/l;off Yaizo,Suruga Bay,Honshu, Japan.

Range; Japan; Australia FC 00 020024

Centroscyllium excelsum SHIRAI & NAKAYA 1990 t/l; Emperor Seamont Chain, N.W.Pacific.

Subgenus PARACENTROSCYLLIUM Alcock 1889

Paracentroscyllium ALCOCK 1889, type Paracentroscyllium ornatum ALCOCK 1889

Paracentroscyllium ornatum ALCOCK 1889 t/l; Bay of Bengal

Range; Bay of Bengal and Arabian Sea.

Genus ACULEOLA De Buen 1959

Aculeola DE BUEN 1959, type Aculeola nigra DE BUEN 1959

Aculeola nigra DE BUEN 1959 t/l; Concon,Chile.

Range; Argentina; Chile; Peru.

Genus MICROSCYLLIUM Shirai & Nakaya 1990

Microscyllium SHIRAI & NAKAYA 1990,type Centroscyllium sheikoi DOLGANEV 1986

Centroscyllium sheikoi DOLGANEV 1986 t/l; N.W.Pacific.

Range;N.W.Pacific.

Genus TRIGONOGNATHUS Mochizuki & Ohe 1990

Trigonognathus MOCHIZUKI & OHE 1990, type Trigonognathus kabeyai MOCHIZUKI & OHE 1990

Trigonognathus kabeyai MOCHIZUKI & OHE 1990 t/l; Japan.

Range;Japan.

Family DALATIIDAE

Genus DALATIAS Rafinesque 1810

Dalatias RAFINESQUE 1810 type, Dalatias sparophagus RAFINESQUE 1810

(unidentifiable). Dalatias nocturnus RAFINESQUE 1810 also listed

Scymnus CUVIER 1817, type Squalus americanus GM 1789 = Squalus licha

BONN. 1788 (Preocc. in Insecta)

Scimnus S.D.W. 1837 (error)

Scymnium CUVIER 1839-43, type Squalus niacensis RISSO 1810

Dalatius AGASSIZ 1845 (emend.)

Scymnorhinus BONAPARTE 1846, type S. lichia BONAPARTE = licha BONNATERRE 1788 ( See Hureau & Monod 1973, for use of this name in preference to Dalatias. They regaard sparophagus as unidentifiable)

Barbarodus GISTEL 1848 (to replace Scymnorhinus)

Pseudoscymnus HERRE 1935 type P. boshuensis HERRE

Le liche BROUSSONET 1780 (Cape Breton, France)

Squalus licha BONN 1788( Le cap Breton, S.France)

Squalus americanus GMELIN 1789

Le Squale liche LAC. 1798 (Cape Breton, France)

Squalus nicaensis RISSO 1810 t/l;Nice, France

? Dalatias sparophagus RAFINESQUE 1810 t/l; Sicily

? Dalatias nocturnus RAFINESQUE 1810 t/l; Sicily

Scymnus vulgaris CLOQUET 1822

Squalus scymnus VOIGT 1832 ( in CUVIER)

? Scymnus aquitanensis DE LA PYLAIE 1835

Scymnorhinus phillippsi WHITLEY 1931 t/l; S.Aus.

Pseudoscymnus boshuensis HERRE 1935 t/l; Boshu, Sagami Bay, Japan.

Scymnorhinus brevipinnis J.L.B. SMITH 1936 t/l; Cape Recife, S. Africa.

Dalatias tachiensis SHEN & TING 1972 t/l; Tachi, Taiwan.

Range;East Atlantic;Irish Atlantic Slope to Rio de Oro. Western Med.

One record for W.Atlantic.Japan. Taiwan. S. Africa. Australia.

FC 00 020002

Genus ISISTIUS Gill 1864

Scymnus QUOY & GAIMARD 1824 (part, non CUVIER 1817), type Scymnus brasiliensis QUOY & GAIMARD 1824

Dalatias GRAY 1851 (part, non RAF. 1810).

Isistius GILL 1864, type Scymnus brasiliensis QUOY & GAIMARD 1824

Leius KNER 1865, type Leius ferox QUOY & GAIMARD 1824 = Scymnus brasiliensis.

Scymnus brasiliensis QUOY & GAIMARD 1824 t/l; Brazil.

Squalus (Scymnus) fulgens BENNETT 1840 t/l;Christmas I.,Tropical Pacific (2.5 degrees S., 163 deg. W.,and 55 deg.N.,110 deg. W. according to Bennett,the second definitely in error as it is a terrestrial location.)

Scymnus (Scymnus) brasiliensis var. torquatus MULLER & HENLE 1841 (no loc. Cape Verde according to Dumeril 1861)

Scymnus (Scymnus) brasiliensis var. unicolor MULLER & HENLE 1841 t/l; Mauritius.

Leius ferox KNER 1865 t/l; Australia.

Range;Tropical Atlantic; Mauritius;Australia;Hawaii.

FC 00 020014

Isistius plutodus GARRICK & SPRINGER 1964 t/l; Gulf of Mexico.

Range;Tropical West Atlantic;Japan.

Isistius labialis MENG, CHU &LI 1985 t/l;S.China Sea.

Range; South China Sea.

Genus EUPROTOMICRUS Gill 1864

Scymnus QUOY & GAIMARD 1824 (part) , type Scymnus bispinatus Q.&G. 1824

Euprotomicrus GILL 1864, type Scymnus (Laemargus) labordii MULLER & HENLE 1839

Leiche laborde QUOY & GAIMARD 1824

Scymnus bispinatus QUOY & GAIMARD 1824 t/l; Mauritius.

Scymnus mauritianus QUOY & GAIMARD 1830 t/l; Mauritius.

Scymnus (Laemargus) labordii MULLER & HENLE 1839 (Mauritius; Bourbon)

Euprotomicrus hyalinus EIGENMANN 1891 t/l; Between San Francisco & Honolulu.

Range; Mauritius; Bourbon I., Indian Ocean;Western Australia; New Zealand;Eastern Pacific; S. Atlantic.

FC 00 020034

Genus SQUALIOLUS Smith & Radcliffe 1912

Squaliolus SMITH & RADCLIFFE 1912, type Squaliolus laticaudus SMITH & RADCLIFFE 1912

Squaliolus laticaudus SMITH & RADCLIFFE 1912 t/l; Batangas Bay, Luzon, Philippines.13 42 N., 120 57 E., in 170 fms.

Squaliolus sarmenti NORONHA 1926 t/l; Madeira

Range;Offshore & oceanic in tropical deep waters worldwide.Recently recorded from Biscay.

Squaliolus aliae TENG 1959 t/l; Taiwan

Range; Taiwan

FC 00 020017

Genus HETEROSCYMNOIDES Fowler 1934

Heteroscymnoides FOWLER 1934, type Heteroscymnoides marleyi FOWLER 1934

Heteroscymnoides marleyi FOWLER 1934 t/l; Point Ocean Beach,Natal.

Heteroscymnus longus FOWLER 1925 ( non TANAKA)

Range;South Africa.

Genus EUPROTOMICROIDES Hulley & Penrith 1966

Euprotomicroides HULLEY & PENRITH 1966 type Euprotomicroides zantedeschia HULLEY & PENRITH 1966

Euprotomicroides zantedeschia HULLEY & PENRITH 1966 t/l; West of Capetown, South Africa.

Range; West of Capetown, South Africa. Second specimen from off Uruguay (1980). In oceanic waters.

Genus MOLLISQUAMA Dolganev 1984

Mollisquama DOLGANEV 1984,type Mollisquama parini DOLGANEV 1984

Mollisquama parini DOLGANEV 1984 t/l; Naska submarine ridge, Eastern Pacific.

Range;Naska submarine ridge, Eastern Pacific.

Family SOMNIOSIDAE

Genus SOMNIOSUS Lesueur 1818

Subgenus SOMNIOSUS Lesueur 1818

Squalus GUNN 1765 (non LINNAEUS) for Squalus carcharias GUNN (non LINNAEUS)

Squalus (Acanthorhinus) BLAINVILLE 1816 (part) for Squalus (Acanthorhinus) norvegianus BLAINVILLE 1816

Somniosus LeSUEUR 1818 type; S. brevipinna LeSUEUR = microcephalus BL. & SCHN. 1801

Scymnus FLEMING 1828 (pt.) for Scymnus borealis FLEMING = borealis SCORESBY

Scymnus (Laemargus) MULLER & HENLE 1837 type; Scymnus borealis FLEMING = Scymnus borealis SCORESBY

Somnolentus SWAINSON 1839

Leiodon WOOD 1846, type; Scymnus echinatum WOOD 1846

Dalatias (Somniosus) GRAY 1851 (non RAFINESQUE) type; Scymnus (Laemargus) borealis MULLER & HENLE

Haa-skierding GUNNERUS 1763 t/l; Trondheim, Norway.

Squalus norvegicus GRONOV. 1763 (Inadmissible work) (Norway)

Squalus carcharias GUNNERUS 1765 (non LINN. = Carcharodon) (Norway)

Haa-kiaerringen ROSTED 1788 (Norway

Squalus microcephalus BL. & SCHN. 1801 t/l; Arctic Seas.

Squalus squatina PALLAS 1814 (non LINN.) t/l; White & Arctic Seas.

Squalus (Acanthorhinus) norvegianus BLAINVILLE 1816 (n. nud)

Squalus (Somniosus) brevipinna LESUEUR 1818 t/l; Massachusetts.

Squalus borealis SCORESBY 1820 t/l; Spitzbergen.

Scymnus gunneri THIENEMANN 1828

Scynmus glacialis FABER 1829 t/l; Iceland

Scymnus micropterus VALENC. 1832 t/l; Seine Estuary

Greenland Shark YARRELL 1836

Leiodon echinatum WOOD 1846 t/l; Portland, Maine.

Squalus norvegicus GRAY 1854 t/l; Bjorn I., Norway.

? Somniosus antarcticus WHITLEY 1939 t/l; MacQuarie I. Sub-antarctic.

Range;Arctic Atlantic;south to North Sea & Portugal in the east & to Cape Cod in the west; S. Africa ; Kerguelen; ? MacQuarie I. Sub-antarctic.

Somniosus pacificus BIGELOW & SCHROEDER 1944 t/l; Sagami Sea, Japan.

Range; North Pacific from S.E. Alaska to Japan. Bering Strait.Occasionally south to Puget Sound & California. Australia.

FC 00 020036

Somniosus sp. (QUERO 1976) [COMPAGNO 1984]

An unnamed long-snouted sp. from the Eastern Atlantic. Specimen in Bocage Museum, Lisbon, destroyed.

Subgenus RHINOSCYMNUS Gill 1865

Rhinoscymnus GILL 1865, type Scymnus rostratus RISSO 1826

Heteroscymnus TANAKA 1912, type Heteroscymnus longus TANAKA 1912

Somniosus (Brevisomniosus) QUERO 1975, type Somniosus (Brevisomniosus) bauchotae QUERO 1975

Scymnus rostratus RISSO 1810 t/l; Nice, France.Mediterranean, deep water.

Heteroscymnus longus TANAKA 1912 t/l; Tokyo Market, Japan.

Somniosus (Brevisomniosus) bauchotae QUERO 1975 t/l; Bay of Biscay,46 50 N., 5 10 W.

Range; Mediterranean, deep water. Bay of Biscay. ?Portugal; Japan.

Genus CENTROSCYMNUS Bocage & Capello 1864

Centroscymnus BOCAGE & CAPELLO 1864, type Centroscymnus coelolepis BOCAGE & CAPELLO 1864

Centroselachus GARMAN 1913, type Centrophorus crepidater BOCAGE & CAPELLO 1864

Proscymnodon FOWLER 1933, type Centrophorus plunketi WAITE 1909

Centroscymnus coelolepis BOCAGE & CAPELLO 1864 t/l; Portugal

Scymnodon melas BIGELOW, SCHROEDER & SPRINGER 1953 t/l; Southern New England Coast.

Centroscymnus macrops CHU,MENG & HU 1982 t/l; South China Sea.

Range; E. North Atlantic,from Iceland to Med. & Cape Verde Is.

W. North Atlantic, from Grand Banks to Nantucket;Australia;? N.Z.

South China Sea.

FC 00 020025

C. CREPIDATER (Bocage & Capello) 1864 Longnose Velvet Dogfish;Deepwater Dogfish

Centrophorus crepidater BOCAGE & CAPELLO 1864 t/l; Portugal

Centrophorus rossi ALCOCK 1898 t/l; Travancore Coast, India in 430 fthms.

Centrophorus jonsonii SAEMUNDSSON 1922 t/l; South coast of Iceland.

Centroscymnus furvescens DE BUEN 1959 t/l;Chile

Range N.E. Atlantic; Eastern South Atlantic;India;New Zealand;S.E.Australia;Chile.

FC 00 020012

Centrophorus (Centroscymnus) coelolepis GUNTHER 1870 ( non BOCAGE & CAPELLO 1864

Centroscymnus cryptacanthus REGAN 1906 t/l; Madeira

Range;N.E. Atlantic.

Centroscymnus macracanthus REGAN 1906 t/l;Straits of Magellan, Argentina.

Etmopterus paessleri LONNBERG 1907 t/l;Straits of Magellan, Argentina.

Range; Straits of Magellan, Argentina.

Centroscymnus owstoni GARMAN 1906 t/l; Sagami Bay, Japan.

Range; Japan; New Zealand; S.E.Australia

FC 00 020019

Centrophorus plunketi WAITE 1909 t/l; New Zealand.

Centrophorus waitei THOMPSON 1930 t/l; off Kaikoura, N.Z. (deep water)

Range; New Zealand;Australia.

FC 00 020013

Genus SCYMNODON Bocage & Capello 1864

Scymnodon BOCAGE & CAPELLO 1864, type Scymnodon ringens BOCAGE & CAPELLO 1864

Zameus JORDAN & FOWLER 1903, type Centrophorus squamulosus GUNTHER 1877

Scymnodon rigens BOCAGE & CAPELLO 1864, Diag. fam. Squal. p.3 extra (lapsus, ringens intended, ringor, to snarl)

Scymnodon ringens BOCAGE & CAPELLO 1864, Proc. zool. soc. London p.263, fig. 5 t/l; Portugal.

Range; N. E. Atlantic.

Centrophorus squamulosus GUNTHER 1877 t/l; Inosima, Japan.

? Centroscymnus obscurus VAILLANT 1888 t/l; "Cotes du Soudan"= Tropical French West Africa. (Smalltooth Velvet Dogfish)

? Scymnodon niger CHU, MENG & HU 1982 t/l; SouthChina Sea.

Range; S. Japan;S. E.Australia;?Tropical East & West Atlantic;S. China Sea.

FC 00 020042

Scymnodon ishiharai YANO & TANAKA 1984 t/l; Japan.

Range;Japan.

Genus SCYMNODALATIAS Garrick 1956

Scymnodalatias GARRICK 1956, type Scymnodon sherwoodi ARCHEY 1921

Scymnodon sherwoodi ARCHEY 1921 t/l; New Brighton Beach, Canterbury,New Zealand.

Range;New Zealand.

Scymnodalatias albicauda TANIUCHI & GARRICK 1986 t/l; S.E. Indian Ocean.

Range; S.E. Indian Ocean

FC 00 020035

S. GARRICKI Kukuev & Konovalenko 1988 Azores Dogfish

Scymnodalatias garricki KUKUEV& KONOVALENKO 1988 t/l; Azores

Range;Azores

Scymnodalatias oligodon KUKUEV & KONOVALENKO 1988 t/l; Eastern Pacific.

Range;Eastern Pacific.

Family OXYNOTIDAE

Genus OXYNOTUS Rafinesque 1810

Oxynotus RAFINESQUE 1810, type Squalus centrina LINN. 1758

Squalus (Centrina) CUVIER 1817, type Squalus centrina LINN. 1758

Centrinus SWAINSON 1838 (emendation)

Kentrines ATHENAEUS c. A.D. 200

Regnard de mer BELON 1551 Hist. nat. poissons marin p.46 bis fig.

Vulpecula BELON 1553 Aquat. p. 63, 64 fig.

Centrina RONDELET 1554, Pisc. p.384, fig.; SALVIANI 1554 Aquat. f. 57, pl 56-57

Galeus centrina GESNER 1558, Aquat. p. 1250

Squalus no. 5 ARTEDI 1738 Ichthyologia, Syn. p.95, Gen .p.67

Galeus sp. no. 7 KLEIN 1742

Squalus centrina LINN. 1758 t/l; Mediterranean.

Le Humantin BROUSSONET 1780

La Mielga CORNIDE 1788

Le squale humantin LACEPEDE 1798

Centrina salviani RISSO 1826

Centrina humantin CLOQUET 1820

Centrina oxynotus SWAINSON 1839

Centrina vulpecula MOREAU 1881

Range;N.E.Atlantic.and Med.

Centrina salviana HUTTON 1890 (non salviani RISSO) t/l; New Zealand

Centrina bruniensis OGILBY 1893 t/l; Bruny I., Tasmania.

Range; Australia; Tasmania; N.Z.

FC 00 021001

Oxynotus paradoxus FRADE 1929 t/l; Morocco.

Range; N.E. Atlantic.

Oxynotus caribbaeus CERVIGON 1961 t/l; N. of Blanquilla I., Venezuela.

Range; Caribbaean

Oxynotus japonicus YANO & MUROFUSHI 1985 t/l; Japan.

Range; Japan

Oxynotus shubnikovi MYAGKOV in GUBANOV et.al. 1986 t/l;

Sp. dub.

Family ECHINORHINIDAE

(Bramble Sharks)

Genus ECHINORHINUS Blainville 1816

Squalus (Echinorhinus) BLAINVILLE 1816, type Squalus spinosus GMELIN 1789 = Squalus brucus BONNATERRE 1788.

Scymnus CUVIER 1817 (part), type Scymnus spinosus CUV. 1817 = Squalus spinosus GMELIN 1789

Goniodus AGASSIZ 1838, type Squalus spinosus GMELIN 1789

Centrophorus SWAINSON 1839 (part, included spinosus) (non MULLER & HENLE 1837)

Echinorhinus (Rubusqualus) WHITLEY 1931, type Echinorhinus (Rubusqualus) mccoyi WHITLEY 1931 = Squalus brucus BONN. 1788

Le Chien De Mer boucle BROUSSONET 1780

Squalus brucus BONNATERRE 1788 t/l;"The Ocean" (North Atlantic)

Squalus spinosus GMELIN 1789 t/l; "In Oceano"

Squale boucle LACEPEDE 1798

Leiche boucle CLOQUET 1816-1830

Spinous shark YARRELL 1839

Echinorhinus obesus A. SMITH 1849 t/l; Cape of Good Hope,South Africa.

Echinorhinus (Rubusqualus) mccoyi WHITLEY 1931 t/l; Portland, Victoria, Australia.(Based on McCoy 1887, said to be separable from Gmelin)

Range; North Sea to Tropical West Africa.Mediterranean;South Africa; Australia; New Zealand; Japan; West Atlantic;Arabian Sea; Eastern Pacific.

FC 00 022001

Echinorhinus cookei PIETSCHMANN 1928 t/l; South coast of Kauai, Hawaii.

Range; Hawaiian Is.;New Zealand; Australia

FC 00 022002

ORDER PRISTIOPHORIFORMES

Genus PRISTIOPHORUS Muller & Henle 1837

Fivegill Sawsharks

Pristiophorus MULLER & HENLE 1837, type Pristis cirratus LA THAM 1794

Pristis cirratus LATHAM 1794 t/l; Port Jackson, New Holland = Australia

Pristis cirrhata BLOCH & SCHNEIDER 1801 (on LATHAM )

Squalus anisodon LACEPEDE 1802 t/l; New Holland

Squalus tentaculatus SHAW 1804 t/l; New Holland

Range;N.S.W. to S.W. Aus.;Philippines; S.Africa

FC 00 023002

Pristiophorus nudipinnis GUNTHER 1870 t/l; Tasmania & New Holland

Pristiophorus owenii GUNTHER 1870 (No loc.) Habitat unknown

Range;Victoria; Tasmania; South Australia.

FC 00 023001

Pristiophorus cirratus RICHARDSON 1846 (non LATHAM)

Pristiophorus japonicus GUNTHER 1870 t/l; Japan.

Range; Japan; Korea.

Pristiophorus schroederi SPRINGER & BULLIS 1960 t/l; off Dog Rocks, Cay Sol Bank,Bahamas.

Range; off Bahamas & S.E. Florida.

Pristiophorus sp. STEHMANN Dwarf Sawshark

Western Indian Ocean

Victoria & N.S.W.,Aus.

FC 00 023003

Slope, Queensland.

FC 00 023004

Genus PLIOTREMA Regan 1906

Pliotrema REGAN 1908, type Pliotrema warreni REGAN 1906

Sixgill Sawsharks

Pliotrema warreni REGAN 1906 t/l; Natal;False Bay, Cape of Good Hope.

Range; South Africa.

Order SQUATINIFORMES

Family SQUATINIDAE

Genus SQUATINA A.M.C. Dumeril 1806

Rhine ARISTOTLE Historia Animalium c. 330 B.C.

Squatina PLINY IX 14 Natural History ( AD 23-79 )

Squatina WOTTON 1552; BELON 1553

Rhina SCHAEFFER 1760 (inadmissible work ) no type

Squatina VALMONT 1768 (inadmissible work )

Rhina KLEIN 1776 (inadmissible work )

Rhina WALBAUM 1792 (on Klein) (inadmissible work )

Squatina RISSO 1810, type Squatina vulgaris RISSO 1810 = Squalus squatina

LINNAEUS

Rhina CUV. 1817, admissible, but preoccupied by Bloch & Schneider.

Squalraia DE LA PYLAIE 1835

Thaumas MUNSTER 1842 (preocc.), type T.alifer MUNSTER 1842

Phorcynis THIOLLIERE 1864, type P. catulina THIOLLIERE 1864

Scaldia LE HON 1871 ( preocc.), type S. biforis LE HON 1871,

Trigonodus WINKLER 1876 preocc.), type T. primus WINKLER 1876

Rhine ARISTOTLE Historia Animalium

Squatina PLINY Natural History IX 14

Squatina WOTTON 1552; BELON 1553.

Squalus squatina LINNAEUS 1758 (Oceano Europaeo)

Rhina SCHAEFFER 1760; KLEIN 1776; WALBAUM 1792 ( on KLEIN)

( Inadmissible works )

Monk or Angel Fish BROOKES 1763

Angel fish (Squalus rondeletii) PENNANT 1769 (Cornwall , England)

Angel fish (Squalus rondeletii) BORLASE 1776

L'Ange BROUSSONET 1780

Le squale ange LACEPEDE 1798

? Squatina angelus A.M.C. DUMERIL 1806

Squatina vulgaris RISSO 1810 t/l; Nice, France

Squatina angelus BLAINVILLE 1816 (n. nud)

Squatina laevis CUVIER 1817 (new name for Squalus squatina LINNAEUS)

? Squatina aculeata CUVIER 1817

Squatina angelus BLAINVILLE 1825

Squatina lewis COUCH 1825 ( lapsus for laevis )

Squatina europaea SWAINSON 1839

? Squalraia acephala DE LA PYLAIE 1835

? Squalraia cervicata DE LA PYLAIE 1835

Monk Fish COUCH 1865

Range; North Sea to Mediterranean.

Squatina dumeril LeSUEUR 1818 t/l; No loc.? New York

Squatina dumerili DeKAY 1842 (New York)

Range; Western Atlantic; S. New England to Florida. Northern Gulf of Mex.;Jamaica.South to Brazil.

? Squalus aculeata LINN. 1758

Squatina aculeata DUMERIL in CUVIER 1829 t/l; off Marseilles, France

? Squatina fimbriata MULLER & HENLE 1841 t/l; Mediterranean

Range; Med. & adjacent Atlantic.

Squatina oculata BONAPARTE 1840 t/l; off Italy.

Range; Mediterranean. East Atlantic, Morocco to Algeria.

Squatina vulgaris MULLER & HENLE 1841 (non RISSO )

Squatina japonica BLEEKER 1858 t/l; Nagasaki, Japan

Range; Japan; Korea.

Squatina californica AYRES 1859 t/l;San Francisco, California

Rhina armata PHILIPPI 1887 t/l; Iquique, Chile.

Rhina philippii GARMAN 1913 t/l; Mejillones I., Peru.

Range; S.E. Alaska to Chile

Rhina squatina McCOY 1879 (non LINN.)

Squatina australis REGAN 1906 t/l; Port Jackson, N.S.W., Australia.

Range; S.E. to S.W.Australia.

FC 00 024001

Squatina nebulosa REGAN 1906 t/l; Japan.

Range;Japan.

Squatina africana REGAN 1908 t/l; Durban Bay, Natal.

Range; Off Natal, S. Africa.

Squatina tergocellata McCULOCH 1914 t/l; Great Australian Bight in 160-200 fthms.

Range; Southern Australia

FC 00 O24002

Rhina argentina MARINI 1930 t/l; N. Argentina

Squatina punctata MARINI 1936 t/l; Argentina

Range; Argentina to Uruguay.

Squatina guggenheim MARINI 1936 t/l; Argentina

Range; Argentina.

Squatina tergocellatoides CHEN 1963 t/l; Taiwan

Range;Taiwan. Known only from single specimen.

Squatina formosa SHEN & TING 1972 t/l; Taiwan

Range;Taiwan. Known only from single specimen.

Squatina occulta VOOREN & DE SILVA 1991 t/l; S. Brazil

Range; S. Brazil

Squatina sp. A LAST & STEVENS 1993

Range;Outer shelf & Upper Slope from Cairns, Queensland to Victoria, Aus.

FC 00 024004

Squatina sp. B LAST & STEVENS 1993

Range;Outer shelf & Upper Slope from Broome to Shark Bay, W. Australia.

FC 00 024005

Range; Sea of Cortez (Gulf of California)

ORDER ORECTOLOBIFORMES

Family PARASCYLLIIDAE

Genus PARASCYLLIUM Gill 1862

Parascyllium GILL 1862,type Hemiscyllium variolatum DUMERIL 1853

Parascyllium (Neoparascyllium) WHITLEY 1939, type Parascyllium multimaculatum SCOTT 1934

Hemiscyllium variolatum DUMERIL 1853 t/l; Tasmania.

Parascyllium nuchalis McCOY 1874 t/l; Port Phillip, Victoria.

Parascyllium nuchale MACLEAY 1881

Range;Victoria; Tasmania; South and Southwest Australia.

FC 00 013004

Parascyllium collare RAMSAY & OGILBY 1888 t/l; Port Jackson. N.S.W. in deep water.

Range;N.S.W.,Victoria;Tasmania.

FC 00 013002

Parascyllium ferrugineum McCULLOCH 1911 t/l;off Port Phillip, Victoria.

Parascyllium multimaculatum SCOTT 1934 t/l; North Tasmania.

Range; Victoria, Aus.

FC 00 013005

Range; Southwest Australia.

FC 00 013018

Genus CIRRHOSCYLLIUM Smith & Radcliffe 1913

Cirrhoscyllium SMITH & RADCLIFFE 1913, type Cirrhoscyllium expolitum SMITH & RADCLIFFE 1913

Zev WHITLEY 1927 (Proposed as a replacement for Cirrhoscyllium SMITH & RADCLIFFE 1913, believed preoccupied by Cirriscyllium Ogilby 1908, substitution not required )

Cirrhoscyllium expolitum SMITH & RADCLIFFE 1913 t/l; 21 33 N., 118 13 E.,in 100 fthms.(China Sea between north Luzon and China)

Range; China Sea between north Luzon and China

Cirrhoscyllium japonicum KAMOHARA 1943 t/l; Mimase, Japan.

Range;Japan.

Cirrhoscyllium formosanum TENG 1959 t/l; Taiwan.

Range;Taiwan.

Family BRACHAELURIDAE

Genus BRACHAELURUS Ogilby1907

Brachaelurus OGILBY 1907, type Chiloscyllium modestum GUNTHER 1871 = Squalus waddi BLOCH & SCHNEIDER 1801

Cirriscyllium OGILBY 1908, type Chiloscyllium modestum GUNTHER 1871 = Squalus waddi BLOCH & SCHNEIDER 1801

Heteroscyllium REGAN 1908, type Brachaelurus colcloughi OGILBY 1908

Brachaelurus was proposed in 1907 by Ogilby for Chiloscyllium modestum GUNTHER ( = Squalus waddi ). He later (1908) transferred his new species, colcloughi to Brachaelurus and proposed Cirriscyllium for modestum.

Cirriscyllium OGILBY 1908 is thus a junior synonym of Brachaelurus OGILBY 1907. Regan in 1908 proposed Heteroscyllium for the species colcloughi .

Cirriscyllium OGILBY does not preoccupy Cirrhoscyllium SMITH & RADCLIFFE 1913 for their species expolitum, thus Whitley's Zev 1927 is an unnecessary substitution.

Squalus waddi BLOCH & SCHNEIDER 1801 t/l; New Holland ( near Sydney )

Chiloscyllium modestum GUNTHER 1871 t/l; Queensland

Chiloscyllium furvum RAMSAY & MACLEAY 1880 t/l; Port Jackson, N.S.W.

Chiloscyllium fuscum PARKER & HASWELL 1897 (No loc.)

Range;N.S.W., Queensland, Aus.

FC 00 013007

Brachaelurus colcloughi OGILBY 1908 t/l; Mud Island, Moreton Bay,Queensland

Range; Queensland

FC 00 013013

Dingerkus (1986) synonymizes colcloughi with waddi.Compagno is inclined to seperate them.

Family ORECTOLOBIDAE

(Wobbegongs; Carpet Sharks)

Genus ORECTOLOBUS Bonaparte 1834

Orectolobus BONAPARTE 1834, type Squalus barbatus GMELIN 1789 ( = Squalus maculatus BONNATERRE 1788)

Chrossorhinus A. SMITH 1837, type Squalus lobatus BLOCH & SCHNEIDER 1801 ( = Squalus maculatus BONNATERRE 1788)

Crossorhinus MULLER & HENLE 1837, type Squalus barbatus GMELIN 1789

( = Squalus maculatus BONNATERRE 1788)

Le Barbu BROUSSONET 1780 (New Holland) Memoir sur les differentes Especes de Chiens de Mer.

Squalus maculatus BONNATERRE 1788 t/l; "La mer du sud"

Squalus barbatus GMELIN 1789 t/l; New Holland ( = Sydney, N.S.W.)

Watt's Shark PHILLIP 1789 ( Sydney Cove, Port Jackson, N.S.W. )

Le squale barbu LACEPEDE 1798

Squalus lobatus BLOCH & SCHNEIDER 1801 t/l; New Holland (on Watt's Shark PHILLIP 1789)

Squalus appendiculatus SHAW & NODDER 1806 (on Watt's Shark PHILLIP 1789)

? Squalus labiatus BLEEKER 1857 (n. nud. )

Range; Queensland south to west Australia.

FC 00 013003

Crossorhinus ornatus DE VIS 1883 t/l;Queensland

Orectolobus devisi OGILBY 1916 t/l; Moreton Bay, Queensland

Range; Queensland south to south Australia.

Orectolobus ornatus halei WHITLEY 1940 t/l; South Australia

Range; South Australia

FC 00 013001 (Whole species)

Crossorhinus barbatus MULLER & HENLE 1841 (non GMELIN) (Japan)

Crossorhinus lobatus MULLER & HENLE 1841 (non BLOCH & SCHNEIDER) (Japan)

Orectolobus japonicus REGAN 1906 t/l; Japan.

? Squalus labiatus BLEEKER 1857 (n. nud.) (Japan)

Range; Japan; China; Philippines.

Orectolobus wardi WHITLEY 1939 t/l; Cape Keith, Melville I., North Austrlia.

Range;North Austrlia.

FC 00 013017

Orectolobus sp. HUTCHINS & SWAINSTON 1986 t/l; Coastal Western Australia

FC 00 013016

Genus EUCROSSORHINUS Regan 1908

Eucrossorhinus REGAN 1908, type Crossorhinus dasypogon BLEEKER 1867

Crossorhinus dasypogon BLEEKER 1867 t/l; off Waigiu I.,Indonesia.

Orectolobus ogilbyi REGAN 1909 t/l; Torres Strait, between Australia & New Guinea.

Range;Indonesia.

FC 00 013011

Genus SUTORECTUS Whitley 1939

Sutorectus WHITLEY 1939, type Crossorhinus tentaculatus PETERS 1864

Crossorhinus tentaculatus PETERS 1864 t/l; Adelaide, South Australia.

Range;Queensland; South Australia.

FC 00 013012

Family HEMISCYLLIIDAE

Genus HEMISCYLLIUM Mueller & Henle in A. Smith 1837

Hemiscyllium MULLER & HENLE in A. SMITH 1837, type Squalus ocellatus BONNATERRE 1788

L' Oeille BROUSSONET 1780 (New Holland)

Squalus ocellatus BONNATERRE 1788 t/l; "La mer du Sud" ( On L' Oeille BROUSSONET 1780)

Squalus oculatus GRAY 1827 (Australia)

Range; New Guinea and N.W.Australia to N.S.W.,Australia .?Melanesia.

FC 00 013014

Scyllium freycineti CUVIER in QUOY & GAIMARD 1824 t/l; Waigiu,Indonesia

Scyllium malaisianus LESSON 1824

Scyllium melaisianum LESSON 1830 t/l; Waigiu,Indonesia

Chiloscyllium malaianum MULLER & HENLE 1839

Scyllium malaisianum GUNTHER 1870

? Hemiscyllium malayanum BLEEKER 1854-55 t/l; Batjan and Amboina, Moluccas.

Range;New Guinea and nearby islands.

Hemiscyllium trispeculare RICHARDSON 1843 t/l; Turtle I., Gulf of Carpentaria, Aus.

Hemiscyllium tricuspidare ANON. (lapsus)

Range; Northern Australia;Queensland;? Celebes,Indonesia.

FC 00 013015

Hemiscyllium hallstromi WHITLEY 1967 t/l; Papua New Guinea.

Range; Papua New Guinea.

Hemiscyllium strahani WHITLEY 1967 t/l; New Guinea.

Range; New Guinea.

Genus CHILOSCYLLIUM Muller & Henle 1837

Chiloscyllium MULLER & HENLE 1837, type Scyllium plagiosum BENNETT 1830

Synchismus GILL 1862, type Squalus tuberculatus BLOCH &SCHNEIDER 1801 = Squalus indicus GMELIN 1789

Chiloscyllium valid as a subgenus for plagiosum; griseum; punctatum

Synchismus valid as a subgenus for indicum.

Squalus sp. GRONOV. 1754

? Squalus colax MEUSCHEN 1781 Index Gronow's Zoophylacium,Pisces 1781 (Name unavailable)

Squalus indicus GMELIN 1789 t/l; Indian Ocean

Le Squale dentele LACEPEDE 1798

Le Squale gronovien LACEPEDE 1798 (Les mers de l'Inde)

Squalus tuberculatus BLOCH & SCHNEIDER 1801 (on Le Squale dentele LACEPEDE 1798)

Squalus gronovianus SHAW 1804 (on Le Squale gronovien LACEPEDE 1798) Indian Seas.

? Squalus (Scyliorhinus) variegatus BLAINVILLE 1816 (n. nud.)

? Squalus (Scyliorhinus) dentatus BLAINVILLE 1816 (n. nud.)

? Squalus (Scyliorhinus) lambarda BLAINVILLE 1816 (n. nud.)

Chiloscyllium phymatodes BLEEKER 1852 t/l; Samarang, Indonesia.

Squalus caudatus GRAY 1854 ( "In mari indico")

Chiloscyllium indicim var. [delta, epsilon, zeta] GUNTHER 1870

Chiloscyllium (Synchismus) colax WHITLEY 1934

Range; Red Sea south to South Africa and east to Taiwan,Philippines and Melanesia.

C. plagiosum plagiosum (BENNETT) 1830 Whitespotted Bambooshark

Scyllium plagiosum BENNETT 1830 t/l; Sumatra

Scyllium ornatum GRAY 1832

Chiloscyllium plagiosum var. interruptum BLEEKER 1852

Chiloscyllium indicum var. obscura GUNTHER 1870 (part, non GRAY 1851) (India;Cape Seas; Ceylon;China)

Range; India to Indonesia; Japan; Philippines.

C. plagiosum caeruleopunctatum PELLEGRIN 1914 Bluespotted Bambooshark

Chiloscyllium caeruleopunctatum PELLEGRIN 1914 t/l; Fort Dauphin, Madagascar.

Range; Madagascar.

Species Range; India to Indonesia; Japan; Philippines; Madagascar.

? Squalus (Scyliorhinus) unicolor BLAINVILLE 1816 ( n. nud.)

? Scyllium griseum VAN HASSELT 1823 (n. nud.) t/l; Java

Chiloscyllium griseum MULLER & HENLE 1841 t/l; Japan (text only)

Chiloscyllium obscurum GRAY 1851 (n. nud.) t/l; Moluccas.

Range; Red Sea to South Africa and east to China and Japan.

? Bokee sorrah RUSSELL 1803 t/l;Vizagapatam, India

? Ra sorrah RUSSELL 1803 t/l;Vizagapatam, India

? Squalus (Scyliorhinus) russellianus BLAINVILLE 1816 (n. nud.)

? Scyllium punctatum VAN HASSELT 1823 ( n. nud.)

Chiloscyllium punctatum MULLER & HENLE 1841 t/l; Java

Chiloscyllium griseum MULLER & HENLE 1841 (plate only)

Chiloscyllium margaritiferum BLEEKER 1854 t/l; Obi I.,Indonesia.

(Dingerkus & DeFino 1983 make margaritiferum BLEEKER 1854 a synonym of plagiosum.

Range; East coast of India; S. Japan;China to Java; Philippines; Southern New Guinea and Northern Australia to Queensland.

FC 00 013008

Chiloscyllium hasseltii BLEEKER 1852 t/l; Batavia, Java

Range; Indonesia.

Chiloscyllium arabicum GUBANOV 1980 t/l; Persian Gulf

Chiloscyllium confusum DINGERKUS & DeFINO 1983 t/l; Arabian Sea.

Range; Persian Gulf ; Arabian Sea.

Chiloscyllium burmensis DINGERKUS & DeFINO 1983 t/l; Bay of Bengal.

Range;Bay of Bengal.

Chiloscyllium dolganovi KHARIN 1983 t/l; Vietnam.

Range; Vietnam.

Family GINGLYMOSTOMATIDAE

Genus GINGLYMOSTOMA Mueller & Henle 1837

Squalus BONNATERRE 1788 (part) ( non LINN.)

Scyllium GRIFFITH in CUVIER 1834 (part) for Scyllium cirrhosum GRIFFITH = Squalus cirratus BONNATERRE 1788

Ginglymostoma MULLER & HENLE 1837, type Squalus cirratus BONNATERRE 1788 by subsequent designation.

Plicodus WINKLER 1874, type Plicodus thielensis WINKLER 1874 (fossil)

Acrodobatis (lapsus) LEIDY 1877, type Acrodobatis serra LEIDY 1877 (fossil)

Acrodontobatis LEIDY 1877 (correction by author, the latter name intended) type Acrodontobatis serra LEIDY 1877 (fossil)

Chien de mer barbillon BROUESSONET 1780

Gata PARRA 1777 (Cuba)

Squalus cirratus BONNATERRE 1788 "American Seas"

Squale barbillon LACEPEDE 1798 (Pacific America)

Squalus punctulatus LACEPEDE 1800 t/l; Cayenne

Squalus punctatus BLOCH & SCHNEIDER 1801

Scyllium cirrhosum GRIFFITH in CUVIER 1834

Squalus argus BANCROFT 1835 t/l; West Indies.

Ginglymostoma fulvum POEY 1861 t/l; Cuba

Ginglymostoma caboverdianus CAPELLO 1867 t/l; Cape Verde Is.

Range; Tropical Atlantic from Carolina to S.Brazil and off tropical West Africa.

In Eastern Pacific,California to Ecuador.

Genus NEBRIUS Ruppell 1835

Nebrius RUPPELL 1835, type Nebrius concolor RUPPELL = Scyllium ferrugineum LESSON

Nebrodes GARMAN 1913 ( substitute name for Nebrius RUPPELL, under the impression that the former was preoccupied by Nebria LATREILLE 1802 [Insecta] and Nebris CUVIER & VALENCIENNES 1830 [Osteichthyes] but this is not so), type Nebrius concolor RUPPELL

Scyllium ferrugineum LESSON 1830 t/l; Port Praslin, New Ireland. Offach Bay,Waigiu.

Nebrius concolor RUPPELL 1835 t/l; Massaua, Red Sea.

Ginglymostoma ruppellii BLEEKER 1852 t/l; Singapore

Ginglymostoma muelleri GUNTHER 1870 t/l; India

Scymnus porosus KLUNZINGER 1871 in HEMPRICH & EHRENBERG 1899 t/l;Red Sea.

Nebrodes macrurus GARMAN 1913 t/l; Port Louis Harbour, Mauritius.

Nebrodes concolor ogilbyi WHITLEY 1934 t/l; Darnley I., Queensland

Nebrius doldi J.L.B. SMITH 1953 t/l; Aldabra I., Western Indian Ocean.

Range;Red Sea to South Africa east to Southern Japan and south to Queensland, Melanesia, Polynesia.

FC 00 013010

Genus PSEUDOGINGLYMOSTOMA Dingerkus 1986

Pseudoginglymostoma DINGERKUS 1986, type Ginglymostoma brevicaudatum GUNTHER in PLAYFAIR & GUNTHER 1866

Ginglymostoma brevicaudatum GUNTHER in PLAYFAIR & GUNTHER 1866

t/l; Zanzibar.

Range; Zanzibar; Seychelles.

Family STEGOSTOMATIIDAE

Genus STEGOSTOMA Mueller & Henle 1837

Stegostoma MULLER & HENLE 1837 type; Squalus fasciatus HERMANN 1783

Squalus sp. GRONOV. 1754

Squalus varius SEBA 1758 Thesauri vol. 3,p. 105. No locality. (Inadmissible work)

Squalus tigrinus PENNANT 1764 (Inadmissible work) t/l; Indian Ocean

Le tigre BROUSSONET 1780

Squalus tigrinus FORSTER 1781

Squalus fasciatus HERMANN 1783 (on SEBA 1758)

Squalus tygrinus BONN. 1788 "La Mer des Indes" (on BROUSSONET 1780)

Squalus tigrinus GMELIN 1789

Squalus longicaudus GMELIN 1789 (on SEBA 1758 and GRONOV. 1763) No locality.

Squalus maculatus BLOCH & SCHNEIDER 1801 (non BONNATERRE 1788) t/l; "Oceano orientale"

Pollee Makum RUSSELL 1803

Scyllium quinquecarinatum VAN HASSELT 1823 t/l; Java

Scyllium heptagonum RUPPELL 1835 t/l; Djedda, Red Sea

Stegostoma carinatum BLYTH 1847 (India)

Squalus pantherinus KUHL & VAN HASSELT 1823 in BLEEKER 1852 (name in synonomy)

Squalus cirrosus GRAY 1854

Stegostoma varium GARMAN 1913 (East Africa)

Stegostoma tigrinum naucum WHITLEY 1939 (Eastern Australia)

Range; Red Sea south to Madagascar and east to Philippines,Japan and N.S.W., Australia. New Caledonia.

FC 00 013006

Family RHINCODONTIDAE

Genus RHINCODON A. Smith 1829

Rhincodon A. SMITH 1829, type Rhincodon typus A. SMITH 1829

Rineodon MULLER & HENLE 1838 (variant spelling)

Rhineodon SWAINSON 1839 (variant spelling)

Rhiniodon SWAINSON 1839 (variant spelling)

Rhinodon MULLER & HENLE 1841 (variant spelling)

Rhineodon GRAY 1857 (variant spelling)

Micristodus GILL 1865, type Micristodus punctatus GILL 1865

Rhincodon typus A. SMITH 1829 t/l; Table Bay, South Africa.

Rhinodon typicus MULLER & HENLE 1841

Micristodus punctatus GILL 1865 t/l; Gulf of California.

Rhinodon pentalineatus KISHINOUYE 1891 t/l; Cape Inul, Japan.

Range;Worldwide in tropical & subtropical surface waters.

FC 00 014001

Order LAMNIFORMES

Family ODONTASPIDAE

Genus ODONTASPIS Agassiz 1838

Odontaspis AGASSIZ 1835 (1838), type Carcharias ferox RISSO 1826 = Squalus ferox RISSO 1810

Triglochis MUELLER & HENLE 1837, type Squalus ferox RISSO 1810

Oxytes GIEBEL 1848, type Oxytes obliqua GIEBEL 1848 (fossil)

ICZN Opinion no. 1459 repealed opinion 733,which had reinstated Carcharias RAF 1810 (suppressed 1965) as valid and having priority over Odontaspis AGASSIZ 1838, where the two are considered synonymous.

Squalus ferox RISSO 1810 t/l;Nice, France.

Odontaspis herbsti WHITLEY 1950 t/l; Gabo I.,N.S.W.,Australia

Range; Med. & adj. Atlantic;Australia;N.Z.;Hawaii; S.California;Gulf of California.; Natal; S.Japan; N.Madagascar.

FC 00 008003

Carcharias noronhai MAUL 1955 t/l; Camara de Lobos,Madeira.

Range;Madeira deep water. S. Brazil. North of Madagascar.

Genus EUGOMPHODUS Gill 1862

Carcharias RAFINESQUE 1810, type Carcharias taurus RAF. 1810

Eugomphodus GILL 1862, type Eugomphodus griseus GILL 1862 = Carcharias griseus STORER 1846 = Squalus littoralis MITCHILL ( = Carcharias taurus RAFINESQUE. )

Synodontaspis WHITE 1931, type Carcharias taurus RAF. 1810

Parodontaspis WHITE 1931, type Odontaspis platensis LAHILLE 1928

Carcharias taurus RAF. 1810 t/l; Sicily

Squalus americanus MITCHILL 1815 t/l; New York.

Squalus littoralis LeSUEUR 1817 t/l; New York.

Squalus macrodus MITCHILL 1817 t/l; New York.

Carcharias griseus AYRES 1842 t/l; Long Island

? Odontaspis cinerea RAMSAY & MACLEAY 1880 (n.nud.) t/l; Pt. Jackson,Australia.

Lamna ecarinata HILGENDORF in HEMPRICH & EHRENBERG 1899 t/l; Alexandria, Egypt.

Carcharias arenarius OGILBY 1911 t/l; Moreton Bay, Queensland.

Carcharias owstoni GARMAN 1913 t/l; Japan.

? Squalus lixa LARRANAGA 1923

Odontaspis platensis LAHILLE 1928 t/l; Argentina.

Range; Both sides of Atlantic; Red Sea to East Indies, Japan and Australia.

FC 00 008001

Carcharias tricuspidatus DAY 1878 t/l; Karachi, Pakistan.

Carcharias cuspidatus OGILBY 1888 (lapsus) t/l; NSW.,Aus.

Range; India. Known only from type,now lost. May be syn. of taurus.

Fossil from Middle Eocene, India.

Family PSEUDOCARCHARIIDAE

Genus PSEUDOCARCHARIAS Cadenat 1963

Pseudocarcharias CADENAT 1963, type Carcharias kamoharai MATSUBARA 1936.

Carcharias kamoharai MATSUBARA 1936 t/l; Koti fish market, Japan.

Carcharias yangi TENG 1959 t/l;off Su-ao, Taiwan.

Carcharias (Pseudocarcharias) pelagicus CADENAT 1963 t/l; Guinea Coast

Range;East China Sea. Japan. Guinea.

FC 00 009003

Family MITSUKURINIDAE

Genus MITSUKURINA Jordan 1898

Rhinognathus DAVIS 1887 (Preoccupied), type Rhinognathus lewisii DAVIS 1887 (Fossil)

Scapanorhynchus WOODWARD 1889 (replacement name for Rhinognathus DAVIS 1887)

Mitsukurina JORDAN 1898, type Mitsukurina owstoni JORDAN 1898

Mitsukurina owstoni JORDAN 1898 t/l; Misaki, Japan.

Odontaspis nasutus DE BRAGANZA 1904 t/l; Portugal

Scapanorhynchus jordani HUSSAKOF 1909 t/l; Japan

Scapanorhynchus dofleinii ENGELHARDT 1912 t/l; Japan.

Scapanorhynchus mitsukurii WHITE 1937 (error for owstonii )

Range; Japan; East & West Atlantic; East Africa; South Australia.

FC 00 009002

Family LAMNIDAE

Squalus LINN. 1758 (part) for Squalus carcharias LINN. 1758

Cynocephalus SCHAEFFER 1760 (inadmissible work), (not KLEIN 1742;1776), type Squalus carcharias LINN. 1758

Carcharias CLOQUET 1817 (part) (non RAF. 1810 )

Carcharodon* A. SMITH 1838, type Carcharias lamia RAF. 1810 = Squalus carcharias LINN. (type by monotypy) (See ICZN. Bull. Zool. Nomencl. 1965)

NOT Carcharodon AGASSIZ 1838, type C. smithii AGASSIZ = n. nud.

NOT Carcharodon AGASSIZ 1838, type C. verus CLOQUET 1822, preoccupied by BLAINVILLE.

Carcharocles JORDAN 1923, type Carcharias auriculatus BLAINVILLE (Fossil)

Carcharhinus WHITLEY 1934 ( non BLAINVILLE 1816 )

* The name Carcharodon for the Great White Shark, was proposed by Mueller & Henle in 1838 with a diagnosis but without mention of a particular species. A type species was designated later the same year by Agassiz as Carcharodon smithii,but this is a nomen nudum not used by M.& H., so Carcharodon must be credited to Agassiz with the type Carcharodon verus AGASSIZ 1838 (="Carcharias"of Agassiz 1835 illustration).The specific name verus dates from CLOQUET 1817-22, it's use not being precluded by Blainville 1816, as the latter is a nomen nudum and therefore has no standing.In 1965, the ICZN ruled that Carcharodon should date from A.Smith 1838, who designated Carcharias lamia RAF. 1810 the type, proposed as a substitute for Squalus carcharias LINN., and therefore type by monotypy.

Kharkharias ATHENAEUS AD 200

Lamia RONDELET 1554

Canis marinus GESNER 1558

Canis carcharias ALDROVANDI 1613

Squalus no. 14 ARTEDI 1738

Squalus carcharias LINNAEUS 1758 t/l; "Oceano Europaeo"

White Shark BROOKES 1763

Cane carcharia CETTI 1777 t/l; Sardinia

Carcharias lamia RAFINESQUE 1810 (subst. for Squalus carcharias

LINNAEUS) t/l; Sicily

Carcharias lamia RISSO 1810 (part) t/l;Nice, France.

Carcharias verus CLOQUET 1822 Mediterranean (non BLAINVILLE 1816)

Carcharias rondeletii ST. VINCENT 1829 (Mediterranean)

Carcharias sp. AGASSIZ 1835 ( illus.)

Squalus (Carcharias) vulgaris RICHARDSON 1836 (America)

Carcharodon capensis A. SMITH 1837 t/l; S. Africa

Carcharodon smithii AGASSIZ 1838 (n. nud); BONAPARTE 1839

Carcharodon lamia BONAPARTE 1839

Carcharodon sp. OWEN 1845

Carcharias atwoodi STORER 1848 t/l; Provincetown, Mass.

? Carcharias vorax OWEN 1853

Carcharias maso NORRIS 1898 (non maou LESSON 1830)

White Pointer STEAD 1933

Carcharodon albimors WHITLEY 1939 (Australia)

Great White Shark KEAN 1944

NOT Squalus carcharias PALLAS 1831 (Kamchatka & Bering Sea) (? = Somniosus )

Range; Mediterranean & adjacent Atlantic to English Channel & Cape Verde Is.;Gulf of Guinea;Nova Scotia to N. Gulf of Mexico; Alaska to Baja California; Panama to Chile;Argentina and S. Brazil; Japan & East China;Philippines; Australia except tropical north;New Zealand;S. Africa; Hawaiian Is.; Micronesia;Cooler coastal waters.

FC 00 010003

Genus LAMNA Cuvier 1817

Squalus (Lamna) CUVIER 1817, type Squalus cornubicus GMELIN 1789 = nasus BONN. 1788

Lamia RISSO 1826 (non FABRICIUS 1775 in Insecta)

Selanonius FLEMING 1828

Exoles GISTEL 1848

Canis carcharias ALDROVANDI 1613 (non ROND. = Carcharodon)

Porbeagle BORLASE 1758 t/l; Cornwall , England.

Squalus glaucus GUNNER 1763 (non LINNAEUS) t/l; Norway (embryo)

Beaumaris Shark PENNANT 1776 (N. Wales)

Haaebrand ASCANIUS 1777

Le Nez BROUSSONET 1780 (Cornwall, England.)

Touille-Boeuf ou Loutre de mer DUHAMEL 1782

Squalus nasus BONN. 1788 (ref. to PENNANT 1776) (Wales)

Squalus cornubicus GMELIN 1789 (Cornwall, England)

Squalus pennanti WALBAUM 1792 (Inadmissible work)

Squale Long-nez LAC. 1798

Squalus monensis SHAW 1804 (Anglesey, N.Wales)

Squalus cornubiensis PENNANT 1812 (Cornwall,England)

Squalus selanonus WALKER( in LEACH 1814)( Scotland)

Lamie nez CLOQUET 1816-30

Selanonius walkeri FLEMING 1828 t/l; Great Britain

Lamna punctata STORER 1839 t/l; Massachusetts Bay

Oxyrhina daekayi GILL 1861 (east coast USA)

Isuropsis dekayi GILL 1871-2 (emendation)

Lamna philippii PEREZ CANTO 1886 t/l; Chile

Lamna sp. LAHILLE 1928 (Argentina)

Lamna whitleyi PHILLIPPS 1935 t/l; Island Bay, New Zealand

Range; Temperate waters of the N. Atlantic; southern S. America;S. Australia; New Zealand;S. Africa and cooler waters of S.W. Indian Ocean.

FC 00 010004

Lamna ditropis HUBBS & FOLLETT 1947 t/l; La Jolla, California.

Range;North Pacific.

Genus ISURUS Rafinesque 1810

Isurus RAFINESQUE 1810, type Isurus oxyrinchus RAFINESQUE 1810

Oxyrhina AGASSIZ 1835, type Lamna oxyrhina CUVIER in VALENC. (MSS) =

oxyrinchus RAFINESQUE

Oxyrrhina BONAPARTE 1846, type Oxyrhina gomphodon MULLER & HENLE

Plectrostoma GISTEL 1848, to replace Oxyrhina AGASSIZ

Isuropsis GILL 1862, type Oxyrhina glaucus MULLER & HENLE 1841

Anatodus LE HON 1871, type A. agassizii Le HON (fossil)

Lamiostoma GLIKMAN 1864

Cane di mare di Messina SPALLANZANI 1793

Isurus oxyrinchus RAFINESQUE 1810 t/l; Sicily

Isurus spallanzani RAFINESQUE 1810 t/l; Sicily

Squalus (Lamna) cepedii LESSON 1830 (equatorial Atlantic)

Lamna oxyrhina AGASSIZ 1838 in CUVIER & VALENCIENNES ms. (teeth); OWEN 1840

Isurus oxyrynchus SWAINSON 1839 (emendation)

Oxyrhina spallanzanii BONAPARTE 1839 t/l; Italy

Oxyrrhina glauca BONAPARTE 1839 t/l; Italy

Oxyrhina gomphodon MUELLER & HENLE 1841

? Lamna latro OWEN 1853 (teeth)

Isuropsis sp. GOODE 1877 (Bermuda)

Lamna guentheri MURRAY 1884 t/l; Karachi, India. (abnormal specimen with double the normal number of teeth.)

Lamna huidobrii PHILIPPI 1887 t/l; Santiago, Chile.

Isurus oxyrhynchus JORDAN and EVERMANN 1895 (emendation)

Isurus mako WHITLEY 1929 t/l;Pt. Jackson, NSW, Australia.

Isurus bideni PHILLIPPS 1932 t/l; South Africa ( jaws and photo )

Isursus tigris africanus J.L.B. SMITH 1957 t/l; South Africa

Range;Warm waters worldwide. Offshore pelagic.

FC 00 010001

Lamna punctata DEKAY 1842 (non STORER 1839 = Squalus nasus BONN. 1788) t/l; New York

? Carcharias tigris ATWOOD 1865 (Gulf of Mexico; Cape Cod)

Isurus paucus GUITART-MANDAY 1966 t/l; ?Cuba

Isurus alatus GARRICK 1967 t/l; Tropical Indian or Pacific Ocean

?Lamiostoma belyaevi GLIKMAN 1964

Range;Western North Atlantic; Tropical West Africa. Central Pacific.Madagascar.

Family ALOPIIDAE

Genus ALOPIAS Rafinesque 1810

Vulpes PLINY, Natural History Libri IX 68

Alopekias ATHENAEUS (AD 200)

Simia BELON 1553

Vulpecula SALVIANI 1554

Vulpes RONDELET 1554

Cercus J.CAIUS 1570

Galeus KLEIN 1742

Vulpecula VALMONT 1768, type Squalus vulpes GMELIN 1768 (Inadmissible work)

Squalus BONN. 1788 (pt.) (non LINNAEUS), type Squalus vulpinus BONN. 1788

Alopias RAFINESQUE 1810, type Alopias macrourus RAFINESQUE= Squalus

vulpinus BONN. 1788

Galeus RAFINESQUE 1810 (part)

Squalus (Carcharhinus) BLAINVILLE 1816 (pt.),included Squalus (Carcharhinus) vulpes BLAINVILLE = vulpinus BONN. 1788

Alopecias MULLER & HENLE 1837, type Carcharias vulpes CUVIER

1817 = Squalus vulpinus BONN. 1788

Alopius SWAINSON 1838 (lapsus)

Vulpecula GARMAN 1913, type Vulpecula marina VALMONT 1768 (inadmissible)

Alopes VLADYKOV & McKENZIE 1939 (error) for Squalus vulpes GM.

Thalassorhinus MULLER & HENLE 1839, type Thalassorhinus vulpecula MUELLER &

HENLE 1839

Vulpes marina PLINY Libri IX 68

Alopekias ATHENAEUS AD. 200

Simia BELON 1553

Vulpecula SALVIANI 1554

Vulpes RONDELET 1555

Vulpecula marina GESNER 1558

Cercus J.CAIUS 1570

Vulpecula marina ALDROVANDI 1613

Le Renard marin PERRAULT 1699

Squalus no. 8 ARTEDI 1738

Galeus cauda longa KLEIN 1742

Sea Fox BORLASE 1758 (Cornwall, England)

Squalus alopecias GRONOV. 1763 (Inadmissible work)

Vulpecula marina VALMONT 1768 (Inadmissible work)

Renard marin DUHAMEL 1769-82

Long-tailed shark PENNANT 1776 t/l; Great Britain

Squalus vulpinus BONNATERRE 1788 t/l; Mediterranean

Squalus vulpes GM. 1789 (Mediterranean, Scotland, Cornwall)

Squale renard LACEPEDE 1798

Galeus vulpecula RAFINESQUE 1810 t/l;Sicily

Alopias macrourus RAFINESQUE 1810 t/l;Sicily

? Galeus thalassinus VALENCIENNES in MULLER & HENLE 1835

Alopias vulpis SWAINS. 1838-9 (lapsus)

? Thalassorhinus vulpecula MULLER & HENLE 1839

Squalus alopecias GRAY 1854 (on Gronov.)

Alopecias barrae PEREZ-CANTO 1886 t/l;Chile

? Alopecias longimana PHILIPPI 1887 (n. nud.)

Alopecias chilensis PHILIPPI 1901 t/l; Valparaiso, Chile

Alopecias longimana PHILIPPI 1901 t/l;Talcahuano Bay,Chile

Vulpecula marina GARMAN 1913

Alopias caudatus PHILIPPS 1932 t/l;New Zealand

Alopias greyi WHITLEY 1937 t/l;New South Wales, Australia (Green Thresher, a colour variety)

Range;Oceanic and coastal. Virtually cicumglobal in warm seas.

FC 00 012001

Alopecias superciliosus LOWE 1840 t/l; Madeira

Alopias profundus NAKAMURA 1935 t/l; Taiwan ( foetus )

Range; Western Med. & E. Atlantic south to S.W. Africa. Eastern USA.

S.Japan & E. China ; Hawaii; N. & E. Australia.;N.Z.; W. Indian

Ocean;S. Brazil.

FC 00 012002

Alopias pelagicus NAKAMURA 1935 t/l; Taiwan ( foetus )

Range;Japan; Taiwan;Hawaii; SW Pacific;Red Sea; Somalia; S. Africa.

FC 00 012003

Alopias sp. EITNER 1995 Systematics of the genus Alopias with evidence for the existence of an unrecognized species. COPEIA 1995 (3)

Range ; Baja California. Probable new species of thresher shark.

Family CETORHINIDAE

Genus CETORHINUS Blainville 1816

Squalus GUNNERUS 1765 (part) (non LINNAEUS) type; Squalus maximus GUNNERUS 1765

Halsydrus NEIL 1809 (n. nud); FLEMING 1817; 1822; type; Halsydrus pontopiddani FLEMING 1817.

Tetraorcas RAFINESQUE 1810 type; Tetraorcas angiona RAF.1810

Tetnorcas RAFINESQUE 1815 (error or emend.)

Squalus (Cetorhinus) BLAINVILLE 1816, type; Squalus ( Cetorhinus) gunneri BLAINV. 1816,substitution for Squalus gunnerianus BLAINVILLE 1810 = Squalus maximus GUNNER.1765

? Scoliophis ANON. 1817

Squalus (Selache) CUVIER 1817 type; S. maximus CUVIER = Squalus maximus GUNN. 1765

Selanche JAROCKI 1822 (error)

Selachus MINDING 1832

Ceteorhinus AGASSIZ 1846 (error)

Polyprosopus COUCH 1867 type; P. macer COUCH 1867

Hannovera BENEDEN 1871 type; H. aurata BENEDEN (fossil)

Cethorhinus ESCRIBANO 1909 (error)

Scapasaurus MARWICK 1942 (n. nud)

Squalus maximus GUNN 1765 t/1; Trondheim, Norway

Basking Shark PENNANT 1769

Le Tres Grand BROUSSONET 1780

Squalus maxima BONNATERRE 1788

Le Squale Tres Grand LACEPEDE 1798

? Halsydrus pontopiddani FLEMING 1809 (n. nud);1817 t/l; Orkneys

Squalus gunnerianus BLAINVILLE 1810

Squalus homianus BLAINVILLE 1810

Squalus pelegrinus BLAINVILLE 1810

? Tetraorcas angiona RAFINESQUE 1810 t/l; Sicily

Squalus peregrinus BLAINVILLE 1811

Squale pelerin BLAINVILLE 1811

Squalus (Cetorhinus) gunneri BLAINVILLE 1816

Squalus (Cetorhinus) shavianus BLAINVILLE 1816

Scoliophis atlanticus ANON 1817

Squalus selanonus LEACH 1818

Squalus rostratus MACRI 1819( Mediterranean)

Squalus isodus MACRI 1819 (Mediterranean)

Squalus elephas LESUEUR 1822 (New Jersey)

Squalus rhinoceros MITCHILL 1828 (New York)

Squalus rashleighanus COUCH 1838 Cornwall (monstrosity)

Squalus cetaceous GRAY 1854 (Norway)

Polyprosopus macer COUCH 1867 (Plymouth) (monstrosity)

Cetorhinus blainvillii BRITO CAPELLO 1870

Hannovera aurata BENEDEN 1871 (fossil)

Selachus pennanti CORNISH 1885

Cetorhinus maccoyi BARRETT 1933

Cetorhinus maximus forma infanuncula DEINSE & ADRIANI 1953

Cetorhinus maximus normani SICCARDI 1961 t/l; S. Atlantic

Basking Shark; Sunfish; Hoe-mother; Homer; Pelerin; Muldoan

Range;N. Norway to Med.,Iceland west to E. Coast of USA, Alaska to Baja California;Ecuador to Chile and Argentina. S. Australia; New Zealand;Japan and East China.

FC 00 011001

Family MEGACHASMIDAE

Genus MEGACHASMA Taylor, Compagno & Struhsaker 1983

Megachasma pelagios TAYLOR, COMPAGNO & STRUHSAKER 1983 t/l; Off Oahu, Hawaiian Is.,1975.

Records; Oahu 1975; off California (WOOD 1986); off Western Australia (1988). 6 specimens now recorded.

FC 00 009001

Order CARCHARHINIFORMES

Subfamily SCYLIORHININAE

Genus SCYLIORHINUS

Genus PORODERMA

Genus CEPHALOSCYLLIUM

Subfamily GALEINAE

Tribe GALEIN1

Subtribe GALEINA

Genus GALEUS

Genus PARMATURUS

Genus CEPHALURUS

Subtribe HALAELURINA

Genus HALAELURUS

Genus HAPLOBLEPHARUS

Genus HOLOHALAELURUS

Genus ASYMBOLUS

Tribe APRISTURINI

Genus APRISTURUS

Subfamily ATELOMYCTERINAE

Genus ATELOMYCTERUS

Genus AULOHALAELURUS

Subfamily SCHROEDERICHTHYINAE

Genus SCHROEDERICHTHYS

Family SCYLIORHINIDAE

Scyliorhinidae Incertae sedis :-

Genus CANINOA Nardo 1841

Caninoa NARDO 1841, type Caninoa chiereghini NARDO 1841 (based on Squalus barbarus CHIERIGHINI undated manuscript)

Caninotus NARDO 1844, emendation for Caninoa NARDO 1841

Thalassoklephetes GISTEL 1848 (to replace Caninoa NARDO)

Caninoa chiereghini NARDO 1841 on Squalus barbarus CHIEREGHINI (MS) t/l;Venice

A dubious species for which Nardo erected a special subfamily Caninoini, but which has never been seen since and may have been a mutilated or malformed specimen with a single dorsal fin like Pentanchus.

Subfamily SCYLIORHININAE

Genus SCYLIORHINUS Blainville 1816

Skyllion ARISTOTLE Historia Animalium

Scyllium THEODORE GAZA (1400-1475)

Galeus BELON 1551

Canicula RONDELET 1554

Catulus GESNER 1558 type Catulus major GES. 1558

Squalus LINN. 1758 (part), type Squalus canicula LINN. 1758

Catulus VALMONT 1768 (Inadmissible work), type Catulus major vulgaris VALMONT 1768 = Squalus canicula LINN. Catulus is preoccupied by Kniphof 1759 (Insecta)

Galeus RAFINESQUE 1810 (part), type Galeus caniculus RAF. 1819 = Squalus canicula LINN.

Squalus (Scyliorhinus) BLAINVILLE 1816, type Squalus canicula LINN.

Scyllium CUVIER 1817, type Squalus canicula LINN.

Scylliorhinus BLAINVILLE 1825 (emendation)

Catulus A. SMITH 1837, type Squalus canicula LINN.

Scyllium (Alphascyllium) LEIGH-SHARPE 1926, type Squalus canicula LINN.

Scyllium (Betascyllium) LEIGH-SHARPE 1926, type Squalus stellaris LINN.

Leigh-Sharpe's names are " pseudogenera " (now reclassed as subgenera) used only within a special scheme for terminology of clasper glans morphology and not available as formal names within the scheme of zoological classification (Jordan 1925).

Scylliorhynchus NOBRE 1935 (error)

Scylliodus AGASSIZ 1843 type S. antiquus AG.

Thyallina AGASSIZ 1843 type T. angusta AG.

Palaeoscyllium WAGNER 1857 type P. decheni MAREK

Trigonodus WINKLER 1876 type T. primus WINKLER

Squalus (Scyliorhinus) myops BLAINVILLE 1816

Squalus (Scyliorhinus) breviculus BLAINVILLE 1816

Squalus (Scyliorhinus) punctatus BLAINVILLE 1816

Squalus (Scyliorhinus) punctulatus BLAINVILLE 1816

Squalus (Scyliorhinus) unicolor BLAINVILLE 1816

Squalus (Scyliorhinus) variegatus BLAINVILLE 1816

Squalus (Scyliorhinus) lambarda BLAINVILLE 1816

May be Scyliorhinids or Orectolobiformes.

S. CANICULUS (Linn.)1758 Lesser Spotted Dogfish; Rough Hound; Sandy Dog Galeus stellaris major BELON 1553

Canicula aristotelis RONDELET 1554

Catulus minor SALVIANI 1554

Kleiner Hundfisch GESNER 1558

Catulus major vulgaris RAY 1713

Squalus sp. ARTEDI 1738

Galeus capito rostroque brevissimus KLEIN 1744

Squalus canicula LINN. 1758

Catulus major vulgaris VALMONT 1768 (Inadmissible work) (part)

Catulus stellaris major DUHAMEL 1769-82

Petite Rousette DUHAMEL 1769-82

Spotted Dogfish (male) PENNANT 1769 (Cornwall)

Lesser Spotted Dogfish (female) PENNANT 1769 (Cornwall)

Le squale rochier LACEPEDE 1798

Galeus catulus RAF. 1810 t/l; Sicily

Squalus (Scyliorhinus) elegans BLAINVILLE 1816 (n.nud.)

Rough Hound COUCH 1867

? Scyllium spinacipellitum VAILLANT 1888.( A doubtful synonym of caniculus according to Compagno (1988), but Seret (1990) made this a senior synonym of Scyliorhinus atlanticus KOEFFED 1927 (Apristurus) and proposed rejection of Vaillant's name in favour of Koeffed.)

? Scyllium acutidens VAILLANT 1888

Scyliorhinus canicula var. albomaculata PIETSCHMANN 1907 t/l; West Coast of Morocco.

Catulus duhamelii GARMAN 1913 (Med.) Dwarf race.

Lesser Spotted Dogfish;Small-Spotted Catshark;Rough Hound;Cur Fish; Daw Fish;Rowhound;Huss;Suss;Sea Nurse;Robin Huss;Morgay; Land Dog; Hondshaie; Kleiner Hundfisch; Petite Rousette.

Range; Norway to Med. & to Black Sea,south to Senegal.

Skyllion asterias ARISTOTLE Historia Animalium

Scyllium stellata THEODORE GAZA (1400-1475)

Galeus stellaris minor BELON 1553

Catulus major SALVIANI 1554

Canicula saxatilis RONDELET 1554

Grosser Hundfisch GESNER 1558

Catulus maximus WILLUGHBY 1686

Squalus stellaris LINN. 1758 t/l; "Oceano Europaeo"

Squalus catulus LINN. 1758

Catulus major vulgaris VALMONT 1768 (Inadmissible work) (part)

Catulus saxatilis VALMONT 1768 ( Inadmissible work)

Le Chat Rochier DUHAMEL 1769-82; BROUSSONET 1780

Greater Spotted Dogfish PENNANT 1769 (Cornwall)

Le squale roussette LACEPEDE 1798

Rock Shark SHAW 1804

Scyllium stellare FLEMING 1828

? Scyllium acanthonotum FILIPPI & VERANY 1853

Nurse Hound COUCH 1867

Range;Atlantic coasts of Europe. Med. to Senegal.

Scyllium capense MULLER & HENLE 1841 t/l; Cape of Good Hope, South Africa.

Scyliorhinus capensis REGAN 1908 (Natal)

Range; South Africa.

S. RETIFER (Garman) 1881 Chain Dogfish; Chain Catshark

Scyllium retiferum GARMAN 1881 t/l; 38 23 N., 73 34 W., off Virginia.

Scyliorhinus retifer JORDAN & GILBERT 1883

Range ; Southern New England to Nicaragua.

Retifer - boa - haeckeli - meadi - besnardi form a sp./subspp. complex.

Scylliorhinus boa GOODE & BEAN 1895 t/l; off Barbados,Lesser Antilles.

Range; Hispaniola; Jamaica; Lesser Antilles; Continental Atlantic coast from Honduras to Colombia. ?Brazil.

Catulus haeckelii RIBEIRO 1907 t/l; Rio de Janeiro,Brazil.

Scyliorhinus fernandezi WEIBEZAHN 1953 t/l; Venezuela

Range; Western Venezuela to Uruguay on Atlantic coast of continental America.

Catulus torazame TANAKA 1908 t/l; Misaki,Sagami Sea, Japan.

Scylliorhinus rudis PIETSCHMANN 1908 t/l; Japan.

Range; Japan; Korea.

Halaelurus garmani FOWLER 1934 t/l; East Indies.

Range; East Indies.

Catulus boae SANCHEZ-ROIG 1931

Scylliorhinus torrei HOWELL-RIVERO 1936 t/l; off Havana, Cuba.

Range; Bahamas; Florida; North coast of Cuba.

Scyliorhinus meadi SPRINGER 1966 t/l; Florida

Range; North Carolina to Florida; Cuba; Bahamas.

Scyliorhinus hesperius SPRINGER 1966 t/l; Caribbaean coast of Panama

Range;Caribbaean coast of Panama,Honduras and Colombia.

Scyliorhinus cervigoni MAURIN & BONNET 1970 t/l; Mauretania

Range; Mauretania to Angola.

Scyliorhinus retifer besnardi SPRINGER & SADOWSKY 1970 t/l; North Uruguay.

Range ; N. Uruguay.

Scyliorhinus comoroensis COMPAGNO 1988 t/l; Comoro Is., Western Indian Ocean.

Range;Comoro Is.

Scyliorhinus tokubee SHIRAI, HAGIWARA & NAKAYA 1992 t/l; Izu Peninsula, Japan.

Range; Izu Peninsula, Japan.

Genus PORODERMA A. Smith 1837

Poroderma A. SMITH 1837, type Squalus africanus GMELIN 1789

Conoporoderma FOWLER 1934, type Scyllium pantherinum MUELLER & HENLE 1841 ( For pantherinus and marleyi )

Le Barbillion BROUSSONET 1780

Le Galonne BROUSSONET 1780

? Squalus fasciatus BONNATERRE 1788

Squalus africanus GMELIN 1789 t/l;" Mari Africanum " Squalus vittatus WALBAUM 1792 (inadmissible work) Name founded on Broussonet's description.

Squalus vittatus SHAW & NODDER 1798

Scyllium africanum A. SMITH 1837 t/l; Cape of Good Hope

Squalus striatus FORSTER in LICHTENSTEIN 1844

Range; South Africa;Mauritius;Madagascar.

Poroderma pantherinum SMITH 1837 (n. nud.)

Scyllium pantherinum SMITH in MULLER & HENLE 1838 t/l; Cape of Good Hope

Poroderma variegatum SMITH 1837 (n. nud.) t/l; Cape of Good Hope

? Poroderma submaculatum SMITH 1837 (n. nud.) t/l; Cape of Good Hope

Scyllium variegatum MULLER & HENLE 1841 t/l; Cape of Good Hope

Scyllium leopardinum (VAN HORST) MULLER & HENLE 1841 (name in synonymy|)Scyllium maeandrinum (RAPP) MULLER & HENLE 1841 (name in synonymy)

Range;Natal, S.Africa;Mauritius

Scyliorhinus regani FOWLER 1925 (non GILCHRIST )

Poroderma marleyi FOWLER 1934 t/l; Natal coast

Scyliorhinus leopardus FOWLER 1935 t/l; Durban, Natal. Lapsus for marleyi

Range;Natal

There are 3 varieties of this species. A "typical" form with rosettes, "marleyi" with large black spots and a "salt and pepper" variety with small black spots.

Genus CEPHALOSCYLLIUM Gill 1862

Cephaloscyllium GILL 186, type Scyllium laticeps DUMERIL 1853

Group 1 Isabellum group

L'Isabelle BROUSSONET 1780

Squalus isabella BONNATERRE 1788 t/l;"La mer du sud"

Squalus sabella GMELIN 1789

Scyllium ? lima RICHARDSON 1842 t/l; New Zealand

Range; New Zealand;Taiwan; SE Honshu,Japan.

Scyllium laticeps DUMERIL 1853 t/l; "New Holland"

Cephaloscyllium indicum JORDAN & SNYDER 1901 (non GMELIN 1789)

Cephaloscyllium laticeps forma nascione WHITLEY 1932 t/l;Montague I., N.S.W., Australia.

Range; South Australia;Tasmania;N.S.W.,Australia.

FC 00 015001

Cephaloscyllium umbratile JORDAN & FOWLER 1903 t/l; Nagasaki, Japan.

Cephaloscyllium formosanum TENG 1962 t/l; Taiwan

Range; Japan; Taiwan

Group 2 Ventiosum group

Scyllium ventriosum GARMAN 1880 t/l; Chile

Catulus uter JORDAN & GILBERT 1896 t/l; California

Range; California to Southern Mexico; Chile.

Group 3 Sufflans group

Scyliorhinus (Cephaloscyllium) sufflans REGAN 1908 t/l; Natal,S. Africa

Range;Natal;S.Mozambique.

Group 4 Fasciatum group

Cephaloscyllium fasciatum CHAN 1966 t/l; Hainan

Range; Hainan;Vietnam;NW. Australia.

FC 00 015007

Group 5 Silasi group

Scyliorhinus (Halaelurus) silasi TALWAR 1974 t/l; off Quilon,Kerala,India.

Range; SW. India.

Un-named Species of Cephaloscyllium

= Cephaloscyllium nascione MAY & MAXWELL 1986 (non WHITLEY) misidentification.

cf. isabella, but not isabella, laticeps or nascione.

Range; N.S.W.; Tasmania;S.Aus.

FC 00 015013

Range; slope off N.E.Aus., between Townsville and Saumarez Reef

FC 00 015030

Range; outer shelf,Northern NSW and Queensland

FC 00 015031

Range; Flinders Rocks, Queensland

FC 00 015032

Range;Rowley Shoals,W.Aus. and Lihou Reef,Queensland

FC 00 015033

Subfamily GALEINAE

Squalus GUNNERUS 1765 (part)

Galeus VALMONT 1768 (Inadmissible work)

Galeus KLEIN 1775 (Inadmissible work)

Galeus RAFINESQUE 1810, type Galeus melastomus RAFINESQUE 1810,by subsequent designation of BONAPARTE 1834.

Scyllium RISSO 1810 (part), (non CUVIER)

Squalus (Scyliorhinus) BLAINVILLE 1825 (part)

Pristiurus BONAPARTE 1831 (nom. nud.)

Scyllium (Pristiurus) BONAPARTE 1834, type Scyllium (Pristiurus) melanostomum BONAPARTE 1834 = Galeus melastomus RAFINESQUE 1810

Pristidurus AGASSIZ 1838 (emendation)

Pristiurus (Figaro) WHITLEY 1928,type Pristiurus ( Figaro )boardmani WHITLEY (subgenus)

Haae gjale STROEM 1762

Squalus catulus GUNNER 1765 (non LINN. 1758) t/l; off Trondhiem,Norway

Rod haae ASCANIUS 1777

Galeus melastomus RAFINESQUE 1810,t/l; Sicily

Squalus catulus (female) RISSO 1810

Squalus (Scyliorhinus) delarochianus BLAINVILLE 1816 (n. nud.);1825

Scyllium artedi RISSO 1820 t/l;Nice, France

Squalus prionurus OTTO 1821

Squalus annulatus NILSSON 1832

Scylliorhinus melanostomus BONAPARTE 1834 t/l; Italy.

Black-mouthed dogfish YARRELL 1836

Pristiurus souverbiei LAFONT 1866 t/l; Coast of Gironde, France

Range; Norway & Faroes to Mediterranean,Morocco & Madeira;South to Senegal.

Pristiurus atlanticus VAILLANT 1888 t/l; Cape Spartal, Morocco.

Range; Morocco & Spain.

Pristiurus murinus COLLETT 1904 t/l;Faroe Channel, off Iceland.

Pristiurus jenseni SAEMUNDSSON 1923 t/l; S.Coast of Iceland.

Range; Iceland;Faroes.

Pristiurus eastmani JORDAN & SNYDER 1904 t/l; off Idzu, Japan.

Range; Japan to Vietnam.

Pristiurus sauteri JORDAN & RICHARDSON 1909 t/l; Takao, Taiwan.

Range; Taiwan. Possibly Japan & Philippines

Pristiurus arae NICHOLLS 1927 t/l; Florida.

Galeus cadenati SPRINGER 1966 t/l; Caribbaean coast of Panama.

Range; Florida to Colombia on the continental coast of America

Galeus arae antillensis SPRINGER 1979 t/l; Leeward Is.

Range; Leeward Is., West Indies

Pristiurus (Figaro) boardmani WHITLEY 1928 t/l; off Montagu I., N.S.W., Australia

Figaro boardmani socius WHITLEY 1939 t/l; Australian Bight.

Range; Western Australia to N.S.W., Australia.

FC 00 015009

Galeus polli CADENAT 1959 t/l; Senegal, West Africa.

Range; Southern Morocco to Namibia.

Galeus piperatus SPRINGER & WAGNER 1966 t/l; Gulf of California

Range; Gulf of California

Galeus nipponensis NAKAYA 1975 t/l; Sea of Japan.

Range; Japan.

Galeus schultzi SPRINGER 1979 t/l; Philippines

Range; Philippines

G. LONGIROSTRIS Tachikawa & Taniuchi 1986 Longnose Sawtail Catshark

Galeus longirostris TACHIKAWA &TANIUCHI 1986 t/l; Japan.

Range; Japan

Galeus sp. A LAST &STEVENS 1993

Galeus gracilis COMPAGNO & STEVENS 1993 t/l; Upper slope ,north of Melville I.,Western Aus. 9 49 S., 135 15 E. in 290 m.

Range; Upper slope W. Australia & Northern Territory, Cape Cuvier to Port Hedland, NW Aus., Melville I., Northern Territory & N. Queensland

Holotype: CSIRO H. 1208

FC 00 015008

N.E. Australia from Rockhampton to Townsville,Queensland.

FC 00 015034

Galeus springeri KONSTANTINOU & COZZI 1998 t/l; Lesser Antilles

Copeia 1998 (1) pp. 151-158

Genus PARMATURUS Garman 1906

Parmaturus GARMAN 1906, type Parmaturus pilosus GARMAN 1906

Dichichthys CHAN 1966, type Dichichthys melanobranchius CHAN 1966

P. XANIURUS (Gilbert) 1892 Filetail Catshark

Catulus xaniurus GILBERT 1892 t/l; Southern & Lower California in 184-684 fms.

Range;Central California to Southern Mexico.

Parmaturus pilosus GARMAN 1906 t/l;Sagami Bay, Japan

Pristiurus hertwigi ENGELHARDT 1912 t/l; Yokohama & Asburatsubo,Japan.

Range; Southern Japan

Dichichthys melanobranchius CHAN 1966 t/l; South China Sea

? Figaro piceus CHU,MENG & LIU 1983 t/l; China.

Range; South China Sea.

Parmaturus campechiensis SPRINGER 1979 t/l; Bay of Campeche, Mexico.

Range;Bay of Campeche, Mexico.

Parmaturus macmillani HARDY 1985 t/l; New Zealand.

Range; N.Z.

Saumarez Plateau off N.E. Australia.

FC 00 015036

Genus CEPHALURUS Bigelow & Schroeder 1941

Cephalurus BIGELOW & SCHROEDER 1941, type Catulus cephalus GILBERT 1891

Catulus cephalus GILBERT 1891 t/l;Gulf of California in 362 fthms. & Clarion Island in 460 fthms.

Range; Gulf of California and Revillagigedo Is.

Subtribe HALAELURINA

Genus HALAELURUS Gill 1862

Subgenus HALAELURUS Gill 1862

Halaelurus GILL 1862, type Scyllium buergeri MUELLER & HENLE 1838

Scyllium (Gammascyllium) LEIGH-SHARPE 1926 type Scyllium buergeri MUELLER & HENLE 1838

Leigh-Sharpe's names are " pseudogenera " (now reclassed as subgenera) used only within a special scheme for terminology of clasper glans morphology and not available as formal names within the scheme of zoological classification (Jordan 1925).

H. BUERGERI (Mueller & Henle) 1838 Blackspotted Catshark

Scyllium buergeri MUELLER & HENLE 1838 t/l; Japan

Range;Japan;Korea;China; Taiwan.

Scyllium quagga ALCOCK 1899 t/l; off Malabar coast,India

Range;Indian Ocean,Malabar and Somalia.

Scyllium natalense REGAN 1904 t/l;Natal

Range; South Africa, Western Cape to Mozambique.

Halaelurus boesemani SPRINGER & D'AUBREY 1972 t/l; off Somalia

Range;Somalia; Gulf of Aden;Vietnam;Philippines;Amboina; N.W.Australia.

FC 00 015004

Halaelurus lineatus BASS,D'AUBREY & KISTNASAMY 1975 t/l;Durban,Natal close inshore.

Range; Natal & S.Mozambique.

Subgenus BYTHAELURUS Compagno 1988

Bythaelurus COMPAGNO 1988, type Scyllium canescens GUNTHER 1988

Scyllium canescens GUNTHER 1878 t/l; S.W. coast of South America.

Range;Peru to Straits of Magellan.

Scyllium hispidum ALCOCK 1891 t/l; Andaman Is.

Range;Andaman Is.,Southern India; ?East Africa.

Scyllium canescens ALCOCK 1896 (non GUNTHER 1878)

Halaelurus alcockii GARMAN 1913 t/l; Arabian Sea.

Range; Arabian Sea, off Pakistan. This is a sp.dub. according to Springer 1979 and Compagno 1984.

Halaelurus dawsoni SPRINGER 1971 t/l; New Zealand

Range; Southern New Zealand

Halaelurus lutarius SPRINGER & D'AUBREY 1972 t/l; Delagoa Bay,Mozambique.

Range;Mozambique;Somalia.

Halaelurus immaculatus CHU & MENG 1982 t/l; South China Sea east of Hainan.

Range; South China Sea east of Hainan.

Halaelurus clevai SERET 1987 t/l; Madgascar

Range;Madgascar

Halaelurus (Bythaelurus) sp. A LAST & STEVENS 1993 Australian Dusky Catshark

Range; Continental slope off Ashmore Reef,North Australia.

FC 00 015035

Haploblepharus GARMAN 1913, type Scyllium edwardsii VOIGT (in CUVIER 1832)

Squalus catulus EDWARDS 1764

Scyllium d'Edwards CUVIER 1817

Scyllium edwardsii VOIGT (in CUVIER 1832)

Range; S. Africa, Cape to Natal.Two forms recognised, a Cape form, and a Natal form,which may be a distinct species, see No.4

Scyllium pictum MUELLER & HENLE 1841 t/l;Cape of Good Hope

Range;S. Africa,Namibia to Cape.

Haploblepharus fuscus J.L.B.Smith 1950 t/l; East London & Knysna, South Africa.

Range;S. Africa,Natal &Cape.

Haploblepharus sp. COMPAGNO Natal Natal Shyshark

Halaelurus (Holohalaelurus) FOWLER 1934, type Scylliorhinus regani GILCHRIST 1922

Scylliorhinus punctatus GILCHRIST & THOMPSON) 1914 t/l; Natal

Scyliorhinus (Halaelurus) polystigma REGAN 1921 t/l; Natal

Range; Natal, South Africa

Scylliorhinus regani GILCHRIST 1922 t/l; Cape Seas.

? Scyliorhinus (Halaelurus) melanostigma NORMAN 1939 t/l; Zanzibar

Range; South Africa. ?Zanzibar

Range; ? Somalia south to Mozambique.

Genus ASYMBOLUS Whitley 1939

Scyliorhinus (Asymbolus) WHITLEY 1939, type Scyllium anale OGILBY 1885

Scyliorhinus (Juncrus) WHITLEY 1939, type Scyllium vincenti ZIETZ 1908

Scyllium maculatum RAMSAY 1880 ( non GUNTHER )

Scyllium anale OGILBY 1885 t/l; Port Jackson, NSW.

Range; NSW. &Tasmania to S.W.Australia.

FC 00 015027

Scyllium vincenti ZIETZ 1908 t/l; Investigator Strait & Kangaroo I.,S. Aus.

Range; NSW. &Tasmania to S.W.Australia.

FC 00 015003

Asymbolus sp .A LAST & STEVENS 1993 Dwarf Catshark Range; NW Australia from Dampier toBuccaneer Archipelago.

FC 00 015022

Range; Recherche Archipelago, S.coast of Western Australia.

FC 00 015023

Asymbolus sp. HUTCHINS & SWAINSTON 1986 S.W.Australia

Range; S.W.Aus.

FC 00 015010

Asymbolus analis LAST,SCOTT &TALBOT 1983 ( non OGILBY ) (misidentification)

Range;Moreton I.,Queensland & Southeast Cape, Tasmania.

FC 00 015024

Range;slope between Swain Reefs & Cairns,NE Australia.

FC 00 015025

Halaelurus analis SCOTT et. al. 1980 (misidentification)

Range; Fowler's Bay, South Australia to Perth, Western Australia. FC 00 015026

Range; New Caledonia

Tribe PENTANCHINI

( APRISTURINI )

Genus APRISTURUS Garman 1913

(Ghost or Demon Catsharks)

Pentanchus SMITH & RADCLIFFE 1912, type Pentanchus profundicolus SMITH & RADCLIFFE 1912 Founded on an abnormal one finned specimen of Apristurus and would in that case be the senior synonym of the latter unless referred to the Official List of Rejected Names in Zoology.

Apristurus GARMAN 1913, type Scylliorhinus indicus BRAUER 1906

Apristurius SCHULZE 1926 (misspelling)

Pentanchus (Parapristurus) FOWLER 1934, type Catulus spongiceps GILBERT 1905

Parmaturus (Compagnoia) SPRINGER 1979, type Parmaturus (Compagnoia) manis SPRINGER 1979

A. BRUNNEUS (Gilbert) 1891 Largenose Brown Catshark

Catulus brunneus GILBERT 1891 t/l; No loc. Perhaps S. California.

? Scylliorhinus saldanha BARNARD 1925 t/l; Saldanha Bay, S.Africa in 500 fms.

Apristurus nasutus DE BUEN 1959 t/l; Valparaiso, Chile.

Apristurus japonicus NAKAYA 1975 t/l; Cape Daito, Honshu, Japan.

? Apristurus F [COMPAGNO] 1979 Inst. f. Seefisch., Hamburg ( Bill Bailey,Lousy and Great Shoal Banks between Ireland & Iceland )

Range; Alaska to S. California. Panama to Central Chile.

Iceland south to Morocco.

SW.Cape, S. Africa.

Honshu, Japan.

A. PROFUNDORUM (Goode & Bean) 1895 Smalleye Catshark

Scylliorhinus profundorum GOODE & BEAN 1895 t/l; Delaware Bay, 39 9 N., 72 3 W., in 816 fthms.

Scyliorhinus microps GILCHRIST 1922 t/l; off Table Bay, S. Africa.

Apristurus pinguis DENG, XIONG & ZHAN 1983 t/l; Ryukiu Is., E.China Sea.

Apristurus federovi DOLGANEV 1985 t/l; Honshu, Japan.

Range; NW Atlantic from Newfoundland to Delaware Bay.

NE Atlantic from Iceland to Scotland

SW. Cape Province, S. Africa.

Japan and East China Sea

A . SPONGICEPS (Gilbert) 1905 Spongehead Catshark

Catulus spongiceps GILBERT 1905 t/l;Bird I., Hawaii.

Range; Hawaiian Is. Nominal record from Celebes.

A. INDICUS (Brauer) 1906 Smallbelly Catshark

Scyliorhinus indicus BRAUER 1906 t/l; 2 58 N., 47 E., Indian Ocean, off Southern Somalia.

Pentanchus (Parapristurus) investigatoris MISRA 1962 t/l; Andaman Sea.

Range; W. Indian Ocean incl. Gulf of Aden and Gulf of Oman.

Andaman Sea.

A. PLATYRHYNCHUS (Tanaka) 1909 Spatulasnout Catshark

Scyliorhinus platyrhynchus TANAKA 1909 t/l; Misaki Sagami, Honshu, Japan.

? Pentanchus sp. JORDAN & HUBBS 1925 Japan.

Pentanchus verweyi FOWLER 1934 t/l; 04 13 N., 118 37 E., Sibuko Bay, Sipadan I., Borneo.

Apristurus parvipinnis SPRINGER & HEEMSTRA 1979 t/l;off Pensacola, Florida

Apristurus canutus SPRINGER & HEEMSTRA 1979 t/l; near Anguilla, Leeward Is.,W. Indies.

Apristurus sinensis CHU & HU 1981 t/l; South China Sea.

Apristurus acanutus MENG,CHU & LI 1985 t/l; South China Sea.

? Apristurus sp. A LAST & STEVENS 1993 FC 00 015014

Range; Japan south to Borneo. ? NW and S.E. Australia.

Florida through West Indies and Gulf of Mexico to Southern Carribbean and Giuanas

A. MACRORHYNCHUS (Tanaka) 1909 Flathead Catshark

Scyliorhinus macrorhynchus TANAKA 1909 t/l; Misaki Sagami, Honshu, Japan.

Scyllium laurussonii SAEMUNDSSON 1922 t/l; S.coast of Iceland.

Apristurus maderensis CADENAT & MAUL 1966 t/l; Madeira

Range; S. Iceland south to Canary Is.

? South Africa

Massachusetts to Northern Gulf of Mexico.

S. Japan; Taiwan

Pentanchus profundicolus SMITH & RADCLIFFE 1912 t/l; 10 02 N., 125 19 E., 1000 m. Mindanao Sea, Philippines.

Known only from the type specimen with a single dorsal fin. Founded on what was almost certainly an abnormal one finned specimen of Apristurus herklotsi .

Pentanchus herklotsi FOWLER 1934 t/l; Cagayan I., Jolo Sea, Philippines

Apristurus longicephalus NAKAYA 1979 t/l; Tosa Bay, Honshu, Japan.

Range; S. Japan; East China Sea; Philippines; NW Australia; Queensland; Seychelles.

FC 00 015021

Apristurus abbreviatus DENG et. al. 1985 t/l; East China Sea.

Apristurus xenolepis MENG,CHU & LI 1985 t/l; South China Sea.

Apristurus brevicaudatus CHU,MENG & LI 1986 t/l; South China Sea.

Apristurus longianaIis CHU,MENG & LI 1986 t/l; South China Sea.

Apristurus longicaudatus CHU,MENG & LI 1986 t/l; South China Sea.

Range; S. Japan to Philippines.

NW Australia; Queensland (FC 00 015021); Seychelles.

A. SIBOGAE (Weber) 1913 Pale Catshark

Scyliorhinus sibogae WEBER 1913 t/l; 00 32 S., 119 39 E., Makassar Str., between Borneo and Celebes, Indonesia in 655 m.

Range; Makassar Str. between Borneo & Celebes.

Scylliorhinus atlanticus KOEFFOED 1927 (non VAILLANT 1888) t/l; Canary Is.

Apristurus riveri BIGELOW & SCHROEDER 1944 t/l; N. coast of Cuba in 580 fms.

Range; Canary Is.

N.coast of Cuba to Western Louisiana.

C. KAMPAE Taylor 1972 Longnose Catshark

Apristurus kampae TAYLOR 1972 t/l; Gulf of California.

Parmaturus (Compagnoia) stenseni SPRINGER 1979 t/l; Gulf of Panama

Parmaturus (Compagnoia) manis SPRINGER 1979 t/l; SW Nantucket, Massachusetts.

Range; Central California to Gulf of California; Gulf of Panama and Panama Bay.

Off Massachusetts; West of Ireland; ?Mauretania.

A number of recently described species have been described from the South China Sea to Australia and SW Pacific, S. Africa and California whose status is still indeterminate in relation to the earlier-named species.

Apristurus gibbosus MENG,CHU & LI 1985 t/l; South China Sea.

Range; South China Sea.

Apristurus macrostomus MENG,CHU & LI 1985 t/l; South China Sea.

Range;South China Sea.

Apristurus micropterygius CHU,MENG & LI 1986 t/l; South China Sea.

Range;South China Sea.

Apristurus sp. SERET New Caledonia

Apristurus sp. SERET Philippines

Apristurus sp. SERET Indonesia

Apristurus sp. COMPAGNO California White-edged Catshark

Apristurus sp. COMPAGNO & EBERT S. Africa Black Wonder Catshark

Apristurus sp. B LAST & STEVENS 1993

Range;slope off Queensland & NSW. & off Geraldton, W. Aus.

FC 00 015015

Apristurus sp. C LAST & STEVENS 1993

Range;NSW. & S. Aus., S. Tasmania, N.Z. 900-1150 m. A possibly allied species occurs well off Perth, W. Aus.

FC 00 015016

Apristurus sp. D LAST & STEVENS 1993

Range;S. Aus., SW. Aus., South Island, N.Z., ? North N.Z.

FC 00 015017

Apristurus sp. E LAST & STEVENS 1993

Range;Broken Bay, NSW. to Beachport, S.Aus.

FC 00 015018

Apristurus sp. F LAST & STEVENS 1993

Range;Perth, W.Aus 3 specimens.

FC 00 015019

Apristurus sp. G LAST & STEVENS 1993

Range; slope from N.Queensland to Tasmania, South & West Aus.Part of a possible complex of species with a N.Z. relative.

FC 00 015020

Apristurus aphyodes NAKAYA & STEHMANN 1998 t/l; George Bligh Bank, N.W. of British Isles

Arch.Fish.Mar.Res. 46(1):77-90

Range; N.E. Atlantic west of British Isles.

Subfamily ATELOMYCTERINAE

Atelomycterus GARMAN 1913, type Scyllium marmoratum BENNETT 1830

Scyllium (Deltascyllium) LEIGH-SHARPE 1926 Leigh-Sharpe's names are " pseudogenera " (now reclassed as subgenera) used only within a special scheme for terminology of clasper glans morphology and not available as formal names within the scheme of zoological classification (Jordan 1925).

Scyllium marmoratum BENNETT 1830 t/l; Sumatra

Scyllium maculatum GRAY in HARDWICKE 1832 (non BLOCH & SCHNEIDER 1801 ) t/l; India

Scyllium ornatum GRAY in HARDWICKE 1832 t/l; China

Scyllium pardum (TEMMINCK) MUELLER & HENLE 1838 (name in synonymy)

Range; Pakistan to Malaya, Philippines,Vietnam and S. China. New Guinea.

Atelomycterus macleayi WHITLEY 1939 t/l; Port Darwin, N.Aus.

Range; Northern Australia.

FC 00 015028

There is currently an unresolved situation in respect of the following two forms;-

Atelomycterus marmoratus McKAY 1966 (non BENNETT) (part) t/l;Western Aus.

Atelomycterus sp.[COMPAGNO] 1984

Atelomycterus sp. SAINSBURY et.al. 1985

Atelomycterus fasciatus COMPAGNO & STEVENS 1993 t/l;Onslow,Western Aus. Holotype: WAM P 8629

Range;

Atelomycteris sp. A LAST & STEVENS 1993 in part Whitespotted Sand Catshark ( ? = fasciatus )

Halaelurus sp. 1 SAINSBURY, LEYLAND & KAILOLA 1985

Atelomycterus macleayi ALLEN & SWAINSTON 1988 (non WHITLEY 1939)

(misidentification)

Atelomycterus sp.[COMPAGNO] 1988

Range; shelf off NW. Australia; Melville I., Gulf of Carpentaria.

FC 00 015005

Genus AULOHALAELURUS Fowler 1934

Halaelurus (Aulohalaelurus) FOWLER 1934, type Catulus labiosus WAITE 1905

? Squalus maculatus BLOCH & SCHNEIDER 1801 (non BONN. 1788) (Oriental Ocean)

Squalus (Scyliorhinus) maculatus BLAINVILLE 1816 (n. nud.)

Scyllium maculatum GUNTHER 1870 (Bramble Bay, Queensland)

Catulus labiosus WAITE 1905 t/l; Freemantle, W. Australia.

Range; Western Australia

FC 00 015029

Aulohalaelurus kanakorum SERET 1990 t/l; New Caledonia

Range;New Caledonia

Subfamily SCHROEDERICHTHYINAE

Genus SCHROEDERICHTHYS Springer 1966

Schroederichthys SPRINGER 1966, type Schroederichthys maculatus SPRINGER 1966

Scyllium bivium MUELLER & HENLE 1841 t/l; "Cape of Good Hope" in error. Does not occur in South African waters.

Scyllium gayi PHILIPPI 1887 t/l; Chile

Range;Chile;Patagonia;Argentina.

Scyllium chilense GUICHENOT 1848 t/l; Chile

? Scyllium brevicolle PHILIPPI 1887 t/l; Valparaiso, Chile

Range;off Peru & Chile.

Schroederichthys maculatus SPRINGER 1966 t/l; Western Caribbean (Range;off Honduras & Nicaragua)

Range;off Honduras & Nicaragua

Schroederichthys tenuis SPRINGER 1966 t/l; Brazil, mouth of the Amazon.

Range;

Family PROSCYLLIIDAE

Genus PROSCYLLIUM Hilgendorf 1904

Proscyllium HILGENDORF 1904, type Scyllium habereri HILGENDORF 1904

Calliscyllium TANAKA 1912, type Calliscyllium venustum TANAKA 1912

Scyllium habereri HILGENDORF 1904 t/l; Takao, Taiwan.

Calliscyllium venustum TANAKA 1912 t/l; Sagami Bay, Japan

Range; S. Japan & Korea suoth to South China Sea. North of Java.

Genus ERIDACNIS H.M. Smith 1913

Eridacnis H.M. SMITH 1913, type Eridacnis radcliffei H.M. SMITH 1913

Neotriakis J.L.B. SMITH 1957, type Neotriakis sinuans J.L.B. SMITH 1957

Eridacnis radcliffei H.M. SMITH 1913 t/l; off Jolo I., Sulu Sea, Philippines

Proscyllium alcocki MISRA 1950 t/l; Andaman Sea

Range;Gulf of Aden;Kenya; Bay of Bengal; S. India; Andamans; Gulf of Tonkin;Philippines.

Triakis barbouri BIGELOW & SCHROEDER 1944 t/l; N.coast of Cuba.

Range; N.coast of Cuba to S.Florida.

Neotriakis sinuans J.L.B. SMITH 1957 t/l; Durban,Natal.

Range; Natal to Mozambique

Genus CTENACIS Compagno 1973

Ctenacis COMPAGNO 1973, type Triakis fehlmanni SPRINGER 1968

Triakis fehlmanni SPRINGER 1968 t/l; SW. of Cape Guardafui,Somalia.

Range; Off Somalia.

FAMILY PSEUDOTRIAKIDAE

Genus PSEUDOTRIAKIS Bocage & Capello 1868

Pseudotriakis CAPELLO 1868, type Pseudotriakis microdon CAPELLO 1868

Pseudotriacis GUNTHER 1870 (Emendation)

Pseudotriakis microdon CAPELLO 1868 t/l; Setubal, Portugal

Pseudotriakis acrales JORDAN & SNYDER 1904 t/l; Japan.

Pseudotriakis acrages GARMAN 1913 (emendation)

Range;N.E.Atlantic;New England;S. Japan;Taiwan; Hawaii; North Madgascar.

Genus GOLLUM Compagno 1973

Gollum COMPAGNO 1973, type Triakis attenuata GARRICK 1954

Triakis attenuata GARRICK 1954 t/l; Cape Palliser, North Island, New Zealand.

Range; New Zealand

Genus and species nov. COMPAGNO, STEHMANN & ANDERSON Pygmy False Catshark

Family LEPTOCHARIIDAE

Genus LEPTOCHARIAS Smith in Muller & Henle 1838

Leptocharias SMITH in MULLER & HENLE 1838, type Triaenodon smithii MULLER & HENLE 1839

Leptocharias MULLER & HENLE 1839 (lapsus)

Leptocarcharias GUNTHER 1870 (emendation)

Trieanodon smithii MULLER & HENLE 1839 t/l; Kabenda Bay,West Africa.

Mustelus osborni FOWLER 1923 t/l; Mouth of the Congo.

Range; E.Atlantic. Mauretania to Angola

Family TRIAKIDAE

Subfamily TRIAKINAE

Tribe TRIAKINI

Genus MUSTELUS Linck 1790

Genus TRIAKIS M.&H. 1838

Tribe SCYLLIOGALEINI

Genus SCYLLIOGALEUS Boulenger 1902

Subfamily GALEORHININAE

Tribe GALEORHININI

Genus GALEORHINUS Blainville 1816

Genus HYPOGALEUS J.L.B.Smith 1957

Tribe IAGINI

Genus HEMITRIAKIS Herre 1923

Genus FURGALEUS Whitley 1951

Genus IAGO Compagno & Springer 1971

Genus GOGOLIA Compagno 1973

Subfamily TRIAKINAE

Genus MUSTELUS Linck 1790

? Galeos ARISTOTLE Historia Animalium

? Mustelus THEODORUS 1470

Mustelus SALVIANI 1554

Mustelus VALMONT 1768 (Inadmissible work), type Galeus stellatus VALMONT 1768 = Squalus mustelus LINN.1758

Mustelus LINCK 1790, type Mustelus laevis LINCK 1790 by monotypy = Squalus mustelus LINN.1758

Galeus RAFINESQUE 1810 (part) Included Squalus mustelus LINN.

Squalus (Galeorhinus) BLAINVILLE 1816 (part), included hinnulus BLAINVILLE = Mustelus asterias CLOQUET, and Squalus galeus LINN. (The tope)

Emissola JAROCKI 1822, type Squalus mustelus FORSTER 1777 (logotype)

Myrmillo GISTEL 1848 ( substitute for Mustelus, believed preoccupied by Mustela LINN.

Rhinotriacis GILL 1862 (part) included Rhinotriacis henlei GILL

Pleuracromylon GILL 1864, type Mustelus laevis RISSO 1826 (n. nud.), Mustelus laevis M.&H. 1841.

Rhinotriakis GILL 1883 (part), included M. laevis RISSO

Cynias GILL 1903, type Squalus canis MITCHILL 1815

Allomycter GUITART 1972, type A. dissutus GUITART 1972 ? = Squalus canis MITCHILL

? Galeos ARISTOTLE Historia Animalium

? Mustelus THEODORUS 1470

Mustelus laevis SALVIANI 1554

Galeus laevis RONDELET 1554

Glatt Seehund GESNER 1558

Squalus mustelus LINN. 1758 t/l; "Oceano Europaeao"

Smooth Dogfish BROOKES 1763

Galeus stellatus VALMONT 1768 (inadmissible work)

Smooth hound PENNANT 1769 (Cornwall, England)

Mustelus laevis LINCK 1790 (substitute for Squalus mustelus LINN.)

Le squale emissole LACEPEDE 1798

Mustelus vulgaris CLOQUET 1821

Mustelus levis RISSO 1826 (emendation)

Mustelus equestris BONAPARTE 1834 (part) Italy

Range; Eastern Atlantic. North Sea to Med. and South Africa.

Squalus canis MTCHILL 1817 t/l; New York

? Allomycter dissutus GUITART 1972

Range; Western Atlantic. Cape Cod to Antilles and Northern Gulf of Mexico. Uruguay and Northern Argentina.

Mustelus canis insularis HEEMSTRA 1997

Mustelus canis insularis HEEMSTRA 1997 t/l; Cay Sal Bank, Bahamas

Range; Bahamas.

Species range; Western Atlantic. Cape Cod to Antilles and Northern Gulf of Mexico.

Uruguay and Northern Argentina.

Galeus asterias RONDELET 1554

Galeus stellatus GESNER 1558

Sternhund GESNER 1558

Mustelus asterias VALMONT 1768 (inadmissible work)

Squalus (Galeorhinus) hinnulus BLAINVILLE 1816 (n. nud.)

Mustelus asterias CLOQUET 1819 (on RONDELET 1554 ) t/l; Europe.

Squalus hinnulus BLAINVILLE 1825

Mustelus stellatus RISSO 1826 t/l; Nice,France

Mustelus plebejus BONAPARTE 1834 t/l; Italy

Mustelus equestris BONAPARTE 1834 (part) Italy

Squalus albomaculatus PLUCAR 1846 t/l; Trieste.

Squalus edentulus DODERLEIN 1881 (non BRUNNICH) t/l; Venice

Range; Eastern North Atlantic. North Sea to Med. and N.W.Africa.

Mustelus punctulatus RISSO 1826 t/l;Nice,France

Mustelus mediterraneus QUIGNARD & CAPAPE 1972 t/l; Mediterranean

Range; Med. & S. Portugal to Morocco.

Mustelus vulgaris SCHLEGEL 1850 (non CLOQUET 1819)

Mustelus manazo BLEEKER 1857 t/l; Nagasaki, Japan.

Range; Japan to Indochina;East Africa; ?Arabian Sea.

Rhinotriacis henlei GILL 1863 t/l; San Francisco, California.

Range; N. California to Gulf of California.Ecuador and Peru.

Mustelus californicus GILL 1864 t/l; San Francisco, California.

Range; N. California to Gulf of California.

Mustelus dorsalis GILL 1864 t/l; Panama.

Range; West Mexico to Peru.

Mustelus antarcticus GUNTHER 1870 t/l; N.S.W.;Tasmania;

Emissola maugeana WHITLEY 1939 t/l; Tasmania

Emissola ganearum WHITLEY 1945 t/l;

Range; Most of Australia except tropical north; Tasmania.

FC 00 017001

Mustelus mento COPE 1877 t/l; Peru

Mustelus edulis PEREZ-CANTO 1886 t/l; Chile

Mustelus abbotti EVERMANN & RADCLIFFE 1917 t/l; Peru.

Range;Chile;Peru;Galapagos;Argentina.

M. LUNULATUS Jordan & Gilbert 1883 Sicklefin Smoothhhound

Mustelus lunulatus JORDAN & GILBERT 1883 t/l; Mazatlan, Mexico.

Range; Southern California to Colombia.

M. MOSIS Hemprich & Ehrenberg 1899 Arabian Hardnose or Moses Smoothhound

Mustelus mosis KLUNZINGER 1871 (n.nud.)

Mustelus mosis HEMPRICH & EHRENBERG 1899 t/l;Red Sea.

Range;Red Sea to India.South Africa.

Mustelus griseus PIETSCHMANN 1908 t/l; Japan.

Cynias kanekonis TANAKA 1916 t/l; Japan.

Range;Japan to Indochina.

Galeorhinus fasciatus GARMAN 1913 t/l; Rio Grande do Sul, Brazil.

Mustelus striatus DEVINCENZI 1920 t/l; Uruguay

Range;Southern Brazil to Northern Argentina.

Mustelus lenticulatus PHILLIPPS 1932 t/l; Wellington Harbour ,New Zealand

Range; New Zealand. Records of this species from Australia are antarcticus.

Mustelus norrisi SPRINGER 1939 t/l; Englewood, Key West, Florida.

Range;Florida to Northern South America on continental coast.

Mustelus schmitti SPRINGER 1939 t/l; off Uruguay and Buenos Aires, Argentina.

Range;S.Brazil to N.Argentina.

Mustelus palumbes J.L.B. SMITH 1957 t/l; Knysna estuary mouth, South Africa.

Range;South Africa.

Mustelus higmani SPRINGER & LOWE 1963 t/l; Surinam

Range;Venezuela to Brazil.

Mustelus whitneyi CHIRICHIGNO 1973 t/l; Peru

Range;Peru to Southern Chile.

Mustelus sp. 4 aff. canis HEEMSTRA 1973; Venezuela

Mustelus minicanis HEEMSTRA 1997 t/l; Gulf of Venezuela, 12 19 N., 70 34 W.

Range; off Venezuela

Mustelus sp. 3 aff. canis HEEMSTRA

Mustelus sinusmexicanus HEEMSTRA 1997 t/l; Dauphin, Alabama

Range; Northern Gulf of Mexico

Mustelus sp. 1 aff. lunulatus HEEMSTRA 1973 Broadnose Smoothhound

Gulf of California

Mustelus sp. 2 aff. lunulatus HEEMSTRA 1973 Pacific Arrownose Smoothhound

Gulf of California

Mustelus sp. 5 HEEMSTRA 1973 Tropical Australia.

Mustelus sp. SAINSBURY,KAILOLA & LEYLAND 1985

Mustelus sp. A LAST & STEVENS 1993

Range; Isolated records from tropical Australia.

FC 00 017005

Mustelus manazo SAINSBURY,KAILOLA & LEYLAND 1985 (non BLEEKER) (Misidentification)

Mustelus sp. STEVENS & McLOUGHLIN 1991

Mustelus sp. LAST & STEVENS 1993

Range; Dampier, West Australia and Cairns, Queensland.

FC 00 017004

Mustelus sp. cf. manazo SERET t/l; New Caledonia.

Genus TRIAKIS Muller & Henle 1837

Triakis MULLER & HENLE 1837, type Triakis scyllium MULLER & HENLE 1841

Triacis GUNTHER 1870 (Emendation)

Genus TRIAKIS Muller & Henle 1837

Triakis MULLER & HENLE 1837, type Triakis scyllium MULLER & HENLE 1841

Triacis GUNTHER 1870 (Emendation)

Triakis scyllium MULLER & HENLE 1841 t/l;Japan.

Hemigaleus pingi EVERMANN & SHAW 1927 t/l; Wenchow, China.

Range; Western North Pacific. Siberia ( Peter the Great Bay ) to China,Japan and Taiwan.

Triakis californica GRAY 1851 (n. nud.)

Triakis semifasciata GIRARD 1854 t/l; San Francisco, California.

Mustelus felis AYRES 1854

Range; Eastern North Pacific from Oregon to Baja California.

Subgenus CAZON De Buen 1959

Cazon DE BUEN 1959, type Triakis maculata KNER & STEINDACHNER 1867

Mustelus megalopterus A. SMITH 1849 t/l; Cape Town.

Mustelus natalensis STEINDACHNER 1866 t/l; Durban, S. Africa.

Mustelus nigropunctatus J.L.B. SMITH 1952 t/l; S. Africa.

Range; Cape Province to Natal, S.Africa.

Triakis scyllium DUMERIL 1865 ( non MULLER & HENLE )

Triakis scyllium var. maculata KNER & STEINDACHNER 1867 t/l;"Mazatlan, Mexico" Original type locality in error, resricted to Callao, Peru by Garman 1913.

Mustelus nigromaculatus EVERMANN& RADCLIFFE 1917 t/l; Lobo de Tierra, Peru.

Range; Peru; Chile;Galapagos.

Triakis acutispina KATO 1968 t/l; Isla de la Plata, Ecuador.

Range; Ecuador.

Tribe SCYLLIOGALEINI

Genus SCYLLIOGALEUS Boulenger 1902

Scylliogaleus BOULENGER 1902, type Scylliogaleus quecketti BOULENGER 1902

Scylliogaleus quecketti BOULENGER 1902 t/l;off Natal, S. Africa.

Range; Natal, S. Africa.

Subfamily GALEORHININAE

? Galeos ARISTOTLE Historia Animalium

Galeus KLEIN 1742 (part)

Galeus SCHAEFFER 1760 (Inadmissible work), type Squalus galeus LINN.

Squalus (Galeorhinus) BLAINVILLE 1816 (part), included hinnulus BLAINVILLE = Mustelus asterias CLOQUET, and Squalus galeus LINN. Type S. galeus LINN. by subsequent designation.

Squalus (Galeus) CUVIER 1817 (non Galeus RAF. 1810)

Eugaleus GILL 1865 type Eugaleus galeus GILL = Squalus galeus LINN.

Notogaleus WHITLEY 1931 type Galeus australis MACLEAY 1881

? Galeos ARISTOTLE Historia Animalium

Galeus canis RONDELET 1554

Canis galeus SALVIANI 1554

Squalus sp. 9 ARTEDI 1738

Squalus galeus LINN. 1758 t/l; Oceano Europaeo

Tope BROOKES 1763

Milandre DUHAMEL 1769-82

? Squalus rhinophanes PETIT & LESUEUR in PERON 1807

Galeus vulgaris FLEMING 1828 t/l; British Isles

Galeus canis BONAPARTE 1834 t/l; Italy

Galeus nilssoni BONAPARTE 1846 t/l; "Northern Ocean"

Galeus communis OWEN 1853

Galeus linnei MALM 1877 t/l; Goteberg, Sweden.

Galeus australis MACLEAY 1881 t/l; Port Jackson, N.S.W., Aus.

Galeus zyopterus JORDAN & GILBERT 1883 t/l; San Pedro, California.

Galeus chilensis PEREZ-CANTO 1886 t/l; Chile

Galeus molinae PHILLIPPI 1887 t/l; Valparaiso, Chile.

Carcharhinus cyrano WHITLEY 1930 t/l; N.S.W.

Galeorhinus vitaminicus DE BUEN 1950 t/l; Lobos I.,Peru.

Range; North Sea to Med. and adjacent Eastern Atlantic. S. and S.W. Africa. Alaska to Lower California. Peru to Southern Brazil. Australia;Tasmania; N.Z.

FC

Genus HYPOGALEUS J.L.B.Smith 1957

Galeorhinus (Hypogaleus) J.L.B SMITH 1957, type Galeorhinus (Hypogaleus) zanzibarensis J.L.B SMITH 1957.

Eugaleus hyugaensis MIYOSI 1939 t/l;Hyuga Nada, Japan.

Galeorhinus (Hypogaleus) zanzibarensis J.L.B SMITH 1957 t/l; Zanzibar

Range;Western Pacific; S.Japan; Taiwan; Australia except Northern coast.

Western Indian Ocean; Tanzania to South Africa.Persian Gulf and Gulf of Oman.

FC 00 017006

Tribe IAGINI

Genus HEMITRIAKIS Herre 1923

Hemitriakis HERRE 1923, type Hemitriakis leucoperiptera HERRE 1923

Galeus japanicus MUELLER & HENLE 1839 t/l; Japan.

Galeorhinus japonicus BLEEKER 1853 (lapsus)

Range; Japan; East China Sea; ?Amboina; ?New Caledonia.

Hemitriakis leucoperiptera HERRE 1923 t/l; Dumaguete,Negros,The Philippines.

Triakis leucoptera FOWLER 1929 (lapsus)

Range; Philippines.

Hemitriakis sp. A LAST & STEVENS 1993

Hemitriakis falcata COMPAGNO & STEVENS 1993 t/l; 16 54 S., 12 50 E., Western Aus.Holotype: WAM P 26271-001

Range; Outer shelf off Port Hedland,W.Aus. 145-200 m. 10 specimens.

FC 00 017009

Hemitriakis sp. B LAST & STEVENS 1993

Hemitriakis abdita COMPAGNO & STEVENS 1993 t/l;17 38 S., 150 11 E.,Coral Sea Plateau, off N.E.Queensland in 225 m. Holotype:CSIRO H 1365 03

Range;Upper slope off Queensland. 6 specimens.

FC 00 017010

Hemitriakis sp. COMPAGNO 1988 Ocellate Topeshark

Hemitriakis sp. (cf. sp. B LAST & STEVENS 1993) New Caledonia

Genus FURGALEUS Whitley 1951

Fur WHITLEY 1943 (preoccupied in Insecta), type Fur macki WHITLEY 1943

Furgaleus WHITLEY 1951, new name for Fur WHITLEY 1943

Fur macki WHITLEY 1943 t/l; Mordiallac,Victoria,Australia

Fur ventralis WHITLEY 1943 t/l;W. & S. Australia.

Range;Tasmania to N.W. Aus.

FC 00 017003

Genus IAGO Compagno & Springer 1971

Iago COMPAGNO & SPRINGER 1971, type Eugaleus omanensis NORMAN 1939

Eugaleus omanensis NORMAN 1939 t/l; Gulf of Oman.

Range; Red Sea and Gulf of Oman to Pakistan.

Iago garricki FOURMANOIR & RIVATON 1979 t/l; Vate,New Hebrides.250-320 m.

Range; New Hebrides.

FC 00 017007

Range; SW.India and Bay of Bengal.

? Triakis lepta ALCOCK (unpublished MS.)

Range; Gulf of Oman & Arabian Sea.

Iago sp. cf. omanensis

Dwarf sp. from Bay of Bengal

Philippines; Western Australia

Genus GOGOLIA Compagno 1973

Gogolia COMPAGNO 1973, type Gogolia filewoodi COMPAGNO 1973

Gogolia filewoodi COMPAGNO 1973 t/l; Gogol River mouth, Astrolabe Bay, New Guinea.

Range; N.E.New Guinea

Family HEMIGALEIDAE

Subfamily HEMIGALEINAE

Genus HEMIGALEUS Bleeker 1852

Hemigaleus BLEEKER 1852, type Hemigaleus microstoma BLEEKER 1852

Negogaleus WHITLEY 1931 (Replacement for Hemigaleus BLEEKER 1852)

Hemigaleus microstoma BLEEKER 1852 t/l; Batavia, Java.

? Hemigaleus machlani HERRE 1929 t/l; Jolo I.,Sulu Sea,Philippines (type destroyed)

? Negogaleus brachygnathus CHU 1960 t/l; China.

Range; Java; S. China; Gulf of Thailand; S. India; N. Australia; ? Philippines.

FC 00 018020

Genus CHAENOGALEUS Gill 1862

Chaenogaleus GILL 1862,type Hemigaleus macrostoma BLEEKER 1852

Hemigaleus macrostoma BLEEKER 1852 t/l; Batavia & Samarang, Java

Hemigaleus balfouri DAY 1878 t/l; Waltair, Coromandel Coast, India.

? Hypoprion notatus BLEEKER 1850 t/l; Java

Range;Persian Gulf to China,Taiwan,Java, Celebes.

Genus PARAGALEUS Budker 1935

Paragaleus BUDKER 1935, type Paragaleus gruveli BUDKER 1935 ( = Hemigaleus pectoralis GARMAN 1906 )

Hemigaleus pectoralis GARMAN 1906 t/l; "Aquarial Gardens", New York, stated to have been collected from Massachusetts and Rhode Island , but the species has never been recorded from the Western Atlantic.

Paragaleus gruveli BUDKER 1935 t/l; off Dakar, Senegal.

Range; Cape Verde Is. and Senegal to Angola.

Negogaleus tengi CHEN 1963 t/l; Taiwan

Negogaleus longicaudatus BESSEDNOV 1966 t/l; Gulf of Tonkin

Range; Taiwan;Vietnam.

Paragaleus leucolomatus COMPAGNO & SMALE 1985 t/l; Kosi Bay, Natal

Range; Natal

Paragaleus sp. aff. pectoralis FOURMANOIR 1961 t/l; Madagascar

Range; Gulf of Oman; NW Indian Ocean.

Genus HEMIPRISTIS Agassiz 1843

Hemipristis AGASSIZ 1843, type Hemipristis serra AGASSIZ 1843 (fossil)

Dirrhizodon KLUNZINGER 1871, type Dirrhizodon elongatus KLUNZINGER 1871

Heterogaleus GOHAR & MAZHAR 1964, type Heterogaleus ghardaquensis GOHAR & MAZHAR 1964 ( = Dirrhizodon elongatus KLUNZINGER 1871)

Dirrhizodon elongatus KLUNZINGER 1871 t/l; Red Sea.

Carcharias ellioti DAY 1878 t/l; Karachi, Pakistan

Hemipristis pingeli SETNA & SARANGDHAR 1946 t/l; Bombay, India.

Paragaleus acutiventralis CHU 1960 t/l; Southern China

Heterogaleus ghardaquensis GOHAR & MAZHAR 1964 t/l;NW. Red Sea.

Range; Red Sea to SW. Cape, S.Africa and east to Vietnam, China & Australia.

FC 00 018011

FAMILY CARCHARHINIDAE

Subfamily GALEOCERDINAE

Genus GALEOCERDO Mueller & Henle 1837

Subfamily SCOLIODONTINAE

Genus SCOLIODON Mueller & Henle 1837

Subfamily CARCHARHININAE

Tribe CARCHARHININI

Genus CARCHARHINUS Blainville 1816

Genus GLYPHIS Agassiz 1843

Genus PRIONACE Cantor 1849

Genus LAMIOPSIS Gill 1862

Genus NEGAPRION Whitley 1940

Genus NASOLAMIA Compagno & Garrick 1983

Tribe RHIZOPRIONODONTINI

Genus RHIZOPRIONODON Whitley 1929

Genus LOXODON Mueller & Henle 1838

Tribe ISOGOMPHODONTINI

Genus ISOGOMPHODON Gill 1862

Tribe TRIAENODONTINI

Genus TRIAENODON Mueller & Henle 1837

Subfamily GALEOCERDINAE

Genus GALEOCERDO Mueller & Henle 1837

Squalus LESUEUR 1822, type Squalus cuvier LESUEUR 1822

Galeus AGASSIZ 1835, type G. cepedianus AGASSIZ (non RAFINESQUE 1810)

Galeocerdo MUELLER & HENLE 1837, type Squalus arcticus FABER 1829 = Squalus cuvier LeSUEUR 1822

Prionodon BLEEKER 1852 (part), type Carcharias (Prionodon) fasciatus BLKR 1852.

Galeodes HECKEL 1854 (preocc), type Galeodes priscus HECKEL (fossil)

Boreogaleus GILL 1861, type Squalus arcticus FABER

? Squalus lamia VALMONT 1768 (inadmissible work)

Canis carcharias DUHAMEL 1769-82 (non ROND., non ALDROVANDI) (inadmissible) (teeth only)

? Le squale milandre LACEPEDE 1789

? Le Squale requin LACEPEDE 1798

Squalus cuvier LESUEUR 1822 t/l;" Terre de Witt, New Holland" ( = N.W.Australia)

Carcharhinus lamia BLAINVILLE 1825 (part)

Squalus arcticus FABER 1829 t/l; Iceland

Galeus cepedianus AGASSIZ 1835 (non PERON 1807; non RAFINESQUE 1810) t/l; E. Indies

Galeus sp. AGASSIZ 1835

? Galeus thalassinus VALENCIENNES in MULLER & HENLE 1835 (see also Alopias vulpinus BONN.)

? Squalus thalassinus MULLER & HENLE 1837 (see also Alopias vulpinus BONN.)

Galeocerdo tigrinus MULLER & HENLE 1839 t/l;India

Galeus maculatus RANZANI 1840 t/l;Brazil

Carcharias (Prionodon) fasciatus BLKR 1852 t/l; Batavia,Java.

Galeocerdo rayneri MCDONALD & BARRON 1868 t/l;Australia

Galeocerdo obtusus KLUNZINGER 1871 (in HEMPRICH & EHRENBERG 1899) t/l; Koseir, Red Sea

Galeocerdo hemprichii KLUNZINGER 1871 (in HEMPRICH & EHRENBERG 1899) t/l; Red Sea

A1 Itachizame MATSUBARA 1955 ( Japan ) ( = G.cuvier )

A2 Sababuki MATSUBARA 1955 ( Japan ) ( = G. rayneri )

Range; All tropical & warm temperate seas.

FC 00 018022

Subfamily SCOLIODONTINAE

Genus SCOLIODON Mueller & Henle 1837

Scoliodon MUELLER & HENLE 1837, type Scoliodon laticaudus MUELLER & HENLE 1838

Physodon MUELLER & HENLE 1839, type Carcharias (Physodon) muelleri MUELLER & HENLE 1839

Alopiopsis LIOY 1865, type Alopiopsis plagiodon LIOY 1865 (fossil)

Pala Sorrah RUSSELL 1803 t/l; Vizagapatam, Madras, India.

Sorra Kowah RUSSELL 1803 t/l; Vizagapatam, Madras, India.

Carcharias palasorrah CUVIER 1829 (on Pala Sorrah RUSSELL 1803)

Carcharias sorrakowah CUVIER 1829 (on Sorra Kowah RUSSELL 1803)

Scoliodon laticaudus MUELLER & HENLE 1838 t/l; India

Carcharias (Physodon) muelleri MUELLER & HENLE 1839 t/l; Bengal

Carcharias (Scoliodon) macrorhynchos BLEEKER 1852 t/l; Batavia, Java

Carcharias (Prionodon) palasorra BLEEKER 1853 t/l; Bengal

Range; Tanzania, East Africa east to China, Japan & Borneo. Close to the continental coast, does not reach Philippines, Red Sea or South Africa.

Subfamily CARCHARHININAE

(Grey Sharks ;Ground Sharks;Whalers; Requins; Galanos)

Nomenclaturally this genus has caused more confusion than any other in the subclass Elasmobranchii. Discussions are to be found in Bigelow & Schroeder (1948), Bass, DAubrey & Kistnasamy (1973), Garrick (1882), Compagno (1984 and 1988) among others. The name was fixed in 1965 by a decision of the I.C.Z.N. to select as neotype a specimen of Carcharias melanopterus QUOY & GAIMARD 1824, as proposed by J.A.F. Garrick in 1962.

Squalus LINN. 1758 (part) Some Squalus carcharias LINN. material may have been this genus but Squalus carcharias LINN. is now universally accepted as the type of Carcharodon A. SMITH.

? Squalus DUHAMEL 1769-1782 (inadmissible work) for Squalus lamia DUHAMEL 1769-1782 ("Nos mers"). Uncertain identity.

? Carcharias RISSO 1810 (part), type Carcharias lamia RISSO 1810

Squalus (Carcharhinus) BLAINVILLE 1816. Blainville included 14 species in his new taxon, all of which are officially nomina nudae :-

glaucus ( now in Prionace CANTOR as presently understood )

caeruleus ( now in Prionace CANTOR as presently understood )

cornubicus ( now in Lamna CUV. as presently understood )

monensis ( now in Lamna CUV. as presently understood )

vulpes ( now in Alopias RAF. as presently understood )

lamia ( a heterogeneous assemblage based on a Carcharhinus species [? longimanus ] and Galeocerdo teeth. )

commersonii ( unidentifiable, may be leucas )

lividus ( unidentifiable, may be leucas )

ustus ( unidentifiable, may be melanopterus )

heterodon ( unidentifiable, may be leucas )

verus ( unidentifiable, may be Carcharodon or leucas)

broussonetii ( unidentifiable , may be leucas )

megalops ( unidentifiable )

heterobranchialis ( unidentifiable )

In 1816, Bosc designated the type as Carcharhinus carcharias but this does not appear in Blainville's 1816 work.

Carcharinus CLOQUET 1817 (emendation)

Squalus (Carcharias) CUVIER 1817 (part) (non RAFINESQUE 1810)

? Squalus (Carcharhinus) BLAINVILLE 1820, type ? Squalus (Carcharhinus) lamia BLAINVILLE 1820 (based on Squalus lamia DUHAMEL 1769-1782 , hence dubious.

Carcharhinus BLAINVILLE in VIEILLOT 1825, type Carcharhinus commersoni BLAINVILLE in VIEILLOT 1825, by subsequent designation of JORDAN & GILBERT 1883, according to Garrick (1982) identifiable as melanopterus Q. & G.

? Carcharias RISSO 1826 (non RAFINESQUE 1810)

Carcharias (Prionodon) MUELLER & HENLE 1839 (part, preoccupied by Horsfield 1823 in Viverridae, Mammalia)

Carcharias (Aprion) MUELLER & HENLE 1839 (preoccupied by Aprion CUVIER & VALENCIENNES 1830 in Osteichthyes), proposed for Carcharias acutidens RUPPELL, and included Carcharias(Aprion) isodon VALENCIENNES in MUELLER & HENLE 1839 & Carcharias(Aprion) brevipinna MUELLER & HENLE 1839. No type designated by MUELLER & HENLE.

Carcharias (Hypoprion) MUELLER & HENLE 1839, type Carcharias (Hypoprion) macloti MUELLER & HENLE 1839

Carcharorhinus AGASSIZ 1846 (?lapsus)

Galeolamna OWEN 1853, type Galeolamna greyi OWEN 1853 (jaws only) = brachyura GUNTH.. Whitley (1940) considered it either brachyurus or macrurus ( = obscurus )

Aprionodon GILL 1862 (new name for Aprion MUELLER & HENLE 1839),type Aprionodon punctatus GILL 1862 ( = Carcharias (Aprion) isodon VALENCIENNES in MUELLER & HENLE 1839, by subsequent designation of Jordan & Evermann 1896). (Not Squalus punctatus MITCHILL 1815, ? = Rhizoprionodon terraenovae RICHARDSON)

Hypoprionodon GILL 1862 (new genus, not an emendation for Hypoprion), type Carcharias (Prionodon) hemiodon MUELLER & HENLE 1839

Eulamia GILL 1862, type Eulamia lamia GILL 1862 (n. nud.) ? = Squalus (Carcharhinus) longimanus POEY ( = maou LESSON )

Platypodon GILL 1862, type Carcharias (Prionodon) menisorrah MUELLER & HENLE ( = falciformis MUELLER & HENLE )

Isoplagiodon GILL 1862, type Carcharias (Prionodon) sorrah MUELLER & HENLE

Gymnorhinus HEMPRICH & EHRENBERG 1899, type Gymnorhinus pharaonis HEMPRICH & EHRENBERG 1899 ( = falciformis MUELLER & HENLE )

Aprionodon GARMAN 1913 (part) included brevipinna and isodon, as well as acutidens ( = Negaprion)

Hypoprion GARMAN 1913 ( included macloti, hemiodon, signatus and playfairii)

Mapolamia WHITLEY 1934, type Carcharias melanopterus QUOY & GAIMARD 1824

Gillisqualus WHITLEY 1934, type Gillisqualus amblyrhynchoides WHITLEY 1934

Galeolamnoides WHITLEY 1934, type Carcharias macrurus RAMSAY & OGILBY 1887 ( = Squalus obscurus LeSUEUR)

Galeolamna (Ogilamia) WHITLEY 1939, type Carcharias stevensi OGILBY 1911 ( = Squalus plumbeus NARDO )

Longmania WHITLEY 1939, type Carcharias (Aprion) brevipinna MUELLER & HENLE

Aprionodon FOWLER 1941 (part), type Carcharias (Aprion) brevipinna MUELLER & HENLE and included acutidens and sitankaiensis ( = falciformis).

Uranga WHITLEY 1943, type Uranga nasuta WHITLEY 1943 ( = Carcharias (Aprion) brevipinna MUELLER & HENLE)

Galeolamna (Uranganops) WHITLEY 1943, type Galeolamna (Uranganops) fitzroyensis WHITLEY 1943

Galeolamna (Lamnarius) WHITLEY 1943, type Galeolamna (Lamnarius) spenceri OGILBY 1911 ( = Carcharias (Prionodon) leucas MUELLER & HENLE

Galeolamna (Bogimba) WHITLEY 1943, type Galeolamna (Bogimba) bogimba WHITLEY 1943 ( = Carcharias [Prionodon] leucas MUELLER & HENLE)

Aprionodon BIGELOW & SCHROEDER 1948 ,type Carcharias (Aprion) brevipinna

Pterolamia SPRINGER 1950 (name preoccupied), type Squalus longimanus POEY 1861 ( = Squalus

(Carcharias) maou LESSON 1830)

Pterolamiops SPRINGER 1951 (new name for Pterolamia SPRINGER )

C. OBSCURUS (Lesueur) 1818 Dusky Shark; Shovelnose Grey Shark; Ridgeback Grey Shark; Common Whaler (Aus.); Black Whaler (Aus.); South Australian Whaler (Aus.);

? Squalus vulpecula SOLANDER 1777 t/l; Australia

Squalus obscurus LESUEUR 1818 t/l; "North America" = East coast of USA.

? Carcharias falcipinnis LOWE 1839 t/l;Madeira

Carcharias (Prionodon) obvelatus VALENCIENNES 1844 t/l; Canary Is.

Carcharias macrurus RAMSAY & OGILBY 1887 t/l; Port Jackson, NSW., Aus.

? Hypoprion / Hemigaleus heterodus PHILIPPI 1887 t/l;Chile

Carcharhinus commersonii LOZANO 1928

Carcharhinus iranzae FOURMANOIR 1961 t/l; Madagascar

Carcharhinus obscurella DENG, XIONG & ZHAN 1981 t/l;S. China

Range; Worldwide in warm temperate & tropical seas, inshore & offshore but not oceanic.

West Atlantic; S.Massachusetts to S. Brazil. Not common between Delaware Bay & N. Florida.

East Atlantic;Madeira; Canaries; Senegal; Cape Verdes;Sierra Leone to Table Bay,S. Africa. Ascension & St. Helena. Natal, Moz., Madagascar; S.Japan; S. China; East & West Australia. Baja California . Formerly thought to be absent from the Mediterranean, there have been some recently confirmed records of this species fron the area.

FC 00 018003

? Squalus (Carcharhinus) ustus BLAINVILLE 1816 (n. nud.)

Carcharias melanopterus QUOY & GAIMARD 1824 t/l; Waigeo I., NW. New Guinea.

Carcharhinus commersonii BLAINVILLE 1825 (as restricted by Garrick 1982)

Squalus ustus DUMERIL 1865

Carcharias (Hypoprion) playfairii GUNTHER 1870 t/l; Zanzibar

Carcharias elegans (KLUNZINGER 1871) HEMPRICH & EHRENBERG 1899 (name in synonymy) t/l; Red Sea

Carcharias marianensis ENGELHARDT 1912 t/l; Guam

Range; Tropical and warm temperate Indo-Pacific from Red Sea to S. Africa east to Japan,Hawaii and Queensland. Recently in the southern Med. via the Suez Canal.

FC 00 018036

Squalus plumbeus NARDO 1827 t/l; Adriatic

Carcharias (Prionodon) milberti VALENCIENNES in MUELLER & HENLE 1841 t/l;New York

Carcharias ceruleus DEKAY 1842 (non Squalus (Carcharhinus) caeruleus BLAINVILLE 1816 = Prionace glauca) t/l; New York to New Hampshire

Lamna caudata DEKAY 1842 t/l; Rhode Island

Squalus caecchia NARDO 1847 t/l; near Venice.

Carcharias (Prionodon) japonicus TEMMINCK & SCHLEGEL 1850 t/l;Japan

? Carcharias leucos GUICHENOT 1850 (non leucas M. & H.) (Algeria)

Carcharias (Prionodon) obtusirostris MOREAU 1881 (Cette, Nice, Atlantic)

Carcharias stevensi OGILBY 1911 t/l; Bustard Bay & N.W.Islet, Queensland, Aus.

Carcharhinus latistomus FANG & WANG 1932 t/l; Tsingtau, China.

Galeolamna (Galeolamnoides) dorsalis WHITLEY 1944 t/l; Western Australia.

Range; Worldwide in tropical & warm temperate seas, common in the Med.

E. Atl; Med.; Tropical W. Africa.

W. Atl; S. New England to S. Brazil.

Red Sea and tropical east Africa. Persian Gulf;Australia; S. Japan to Indochina. Hawaii. Probably absent from East Pacific.

FC 00 018007

Carcharias albimarginatus RUPPELL 1835 t/l; Ras Mahamet, Red Sea

Eulamia (Platypodon) platyrhynchus GILBERT 1892 t/l; Magdalena Bay, Clarion I. off Lower California.

Eulamia albomarginata FOWLER 1929 (lapsus for albimarginata)

Range; Red Sea & Western Indian Ocean; Philippines to Taiwan. Islands of tropical central & eastern Pacific. Not found in Eastern Indian Ocean or Hawaiian Is.

FC 00 018027

? Carcharias fissidens BENNETT 1830 t/l; Atlantic coast of North Africa. It is more probable that this is Rhizoprionodon terraenovae.

Carcharias porosus RANZANI 1939 t/l; Brazil

Carcharias (Prionodon) henlei VALENCIENNES in MUELLER & HENLE 1841 t/l; Cayenne, French Guiana.

Carcharhinus sp. JORDAN 1885

Carcharhinus cerdale GILBERT in JORDAN & EVERMANN 1898 t/l; Pacific coast of Panama.